the project gutenberg ebook the origin species means natural selection this ebook for the use anyone anywhere the united states and most other parts the world cost and with almost restrictions whatsoever you may copy give away use under the terms the project gutenberg license included with this ebook online www gutenberg org you are not located the united states you will have check the laws the country where you are located before using this ebook title the origin species means natural selection author charles darwin release date march 1998 ebook 1228 most recently updated october 2023 language english start the project gutenberg ebook the origin species means natural selection there are several editions this ebook the project gutenberg collection various characteristics each ebook are listed aid selecting the preferred file click any the filenumbers below quickly view each ebook 1228 1859 first edition 22764 1860 second edition 2009 1872 sixth edition considered the definitive edition the origin species means natural selection the preservation favoured races the struggle for life charles darwin fellow the royal geological linnæan etc societies author journal researches during beagle voyage round the world london john murray albemarle street 1859 but with regard the material world can least far this can perceive that events are brought about not insulated interpositions divine power exerted each particular case but the establishment general laws whewell bridgewater treatise conclude therefore let man out weak conceit sobriety ill applied moderation think maintain that man can search too far too well studied the book god word the book god works divinity philosophy but rather let men endeavour endless progress proficience both bacon advancement learning down bromley kent october 1859 contents introduction variation under domestication variation under nature struggle for existence natural selection laws variation difficulties theory instinct hybridism the imperfection the geological record the geological succession organic beings geographical distribution geographical distribution continued mutual affinities organic beings morphology recapitulation and conclusion index deteailed contents the origin species introduction chapter variation under domestication causes variability effects habit correlation growth inheritance character domestic varieties difficulty distinguishing between varieties and species origin domestic varieties from one more species domestic pigeons their differences and origin principle selection anciently followed its effects methodical and unconscious selection unknown origin our domestic productions circumstances favourable man power selection chapter variation under nature variability individual differences doubtful species wide ranging much diffused and common species vary most species the larger genera any country vary more than the species the smaller genera many the species the larger genera resemble varieties being very closely but unequally related each other and having restricted ranges chapter struggle for existence bears natural selection the term used wide sense geometrical powers increase rapid increase naturalised animals and plants nature the checks increase competition universal effects climate protection from the number individuals complex relations all animals and plants throughout nature struggle for life most severe between individuals and varieties the same species often severe between species the same genus the relation organism organism the most important all relations chapter natural selection natural selection its power compared with man selection its power characters trifling importance its power all ages and both sexes sexual selection the generality intercrosses between individuals the same species circumstances favourable and unfavourable natural selection namely intercrossing isolation number individuals slow action extinction caused natural selection divergence character related the diversity inhabitants any small area and naturalisation action natural selection through divergence character and extinction the descendants from common parent explains the grouping all organic beings chapter laws variation effects external conditions use and disuse combined with natural selection organs flight and vision acclimatisation correlation growth compensation and economy growth false correlations multiple rudimentary and lowly organised structures variable parts developed unusual manner are highly variable specific characters more variable than generic secondary sexual characters variable species the same genus vary analogous manner reversions long lost characters summary chapter difficulties theory difficulties the theory descent with modification transitions absence rarity transitional varieties transitions habits life diversified habits the same species species with habits widely different from those their allies organs extreme perfection means transition cases difficulty natura non facit saltum organs small importance organs not all cases absolutely perfect the law unity type and the conditions existence embraced the theory natural selection chapter instinct instincts comparable with habits but different their origin instincts graduated aphides and ants instincts variable domestic instincts their origin natural instincts the cuckoo ostrich and parasitic bees slave making ants hive bee its cell making instinct difficulties the theory the natural selection instincts neuter sterile insects summary chapter hybridism distinction between the sterility first crosses and hybrids sterility various degree not universal affected close interbreeding removed domestication laws governing the sterility hybrids sterility not special endowment but incidental other differences causes the sterility first crosses and hybrids parallelism between the effects changed conditions life and crossing fertility varieties when crossed and their mongrel offspring not universal hybrids and mongrels compared independently their fertility summary chapter the imperfection the geological record the absence intermediate varieties the present day the nature extinct intermediate varieties their number the vast lapse time inferred from the rate deposition and denudation the poorness our palæontological collections the intermittence geological formations the absence intermediate varieties any one formation the sudden appearance groups species their sudden appearance the lowest known fossiliferous strata chapter the geological succession organic beings the slow and successive appearance new species their different rates change species once lost not reappear groups species follow the same general rules their appearance and disappearance single species extinction simultaneous changes the forms life throughout the world the affinities extinct species each other and living species the state development ancient forms the succession the same types within the same areas summary preceding and present chapters chapter geographical distribution present distribution cannot accounted for differences physical conditions importance barriers affinity the productions the same continent centres creation means dispersal changes climate and the level the land and occasional means dispersal during the glacial period extensive with the world chapter geographical distribution continued distribution fresh water productions the inhabitants oceanic islands absence batrachians and terrestrial mammals the relation the inhabitants islands those the nearest mainland colonisation from the nearest source with subsequent modification summary the last and present chapters chapter mutual affinities organic beings morphology embryology rudimentary organs classification groups subordinate groups natural system rules and difficulties classification explained the theory descent with modification classification varieties descent always used classification analogical adaptive characters affinities general complex and radiating extinction separates and defines groups morphology between members the same class between parts the same individual embryology laws explained variations not supervening early age and being inherited corresponding age rudimentary organs their origin explained summary chapter recapitulation and conclusion recapitulation the difficulties the theory natural selection recapitulation the general and special circumstances its favour causes the general belief the immutability species how far the theory natural selection may extended effects its adoption the study natural history concluding remarks the origin species introduction when board beagle naturalist was much struck with certain facts the distribution the inhabitants south america and the geological relations the present the past inhabitants that continent these facts seemed throw some light the origin species that mystery mysteries has been called one our greatest philosophers return home occurred 1837 that something might perhaps made out this question patiently accumulating and reflecting all sorts facts which could possibly have any bearing after five years work allowed myself speculate the subject and drew some short notes these enlarged 1844 into sketch the conclusions which then seemed probable from that period the present day have steadily pursued the same object hope that may excused for entering these personal details give them show that have not been hasty coming decision work now nearly finished but will take two three more years complete and health far from strong have been urged publish this abstract have more especially been induced this wallace who now studying the natural history the malay archipelago has arrived almost exactly the same general conclusions that have the origin species last year sent memoir this subject with request that would forward sir charles lyell who sent the linnean society and published the third volume the journal that society sir lyell and hooker who both knew work the latter having read sketch 1844 honoured thinking advisable publish with wallace excellent memoir some brief extracts from manuscripts this abstract which now publish must necessarily imperfect cannot here give references and authorities for several statements and must trust the reader reposing some confidence accuracy doubt errors will have crept though hope have always been cautious trusting good authorities alone can here give only the general conclusions which have arrived with few facts illustration but which hope most cases will suffice one can feel more sensible than the necessity hereafter publishing detail all the facts with references which conclusions have been grounded and hope future work this for well aware that scarcely single point discussed this volume which facts cannot adduced often apparently leading conclusions directly opposite those which have arrived fair result can obtained only fully stating and balancing the facts and arguments both sides each question and this cannot possibly here done much regret that want space prevents having the satisfaction acknowledging the generous assistance which have received from very many naturalists some them personally unknown cannot however let this opportunity pass without expressing deep obligations hooker who for the last fifteen years has aided every possible way his large stores knowledge and his excellent judgment considering the origin species quite conceivable that naturalist reflecting the mutual affinities organic beings their embryological relations their geographical distribution geological succession and other such facts might come the conclusion that each species had not been independently created but had descended like varieties from other species nevertheless such conclusion even well founded would unsatisfactory until could shown how the innumerable species inhabiting this world have been modified acquire that perfection structure and coadaptation which most justly excites our admiration naturalists continually refer external conditions such climate food etc the only possible cause variation one very limited sense shall hereafter see this may true but preposterous attribute mere external conditions the structure for instance the woodpecker with its feet tail beak and tongue admirably adapted catch insects under the bark trees the case the misseltoe which draws its nourishment from certain trees which has seeds that must transported certain birds and which has flowers with separate sexes absolutely requiring the agency certain insects bring pollen from one flower the other equally preposterous account for the structure this parasite with its relations several distinct organic beings the effects external conditions habit the volition the plant itself the author the vestiges creation would presume say that after certain unknown number generations some bird had given birth woodpecker and some plant the misseltoe and that these had been produced perfect now see them but this assumption seems explanation for leaves the case the coadaptations organic beings each other and their physical conditions life untouched and unexplained therefore the highest importance gain clear insight into the means modification and coadaptation the commencement observations seemed probable that careful study domesticated animals and cultivated plants would offer the best chance making out this obscure problem nor have been disappointed this and all other perplexing cases have invariably found that our knowledge imperfect though variation under domestication afforded the best and safest clue may venture express conviction the high value such studies although they have been very commonly neglected naturalists from these considerations shall devote the first chapter this abstract variation under domestication shall thus see that large amount hereditary modification least possible and what equally more important shall see how great the power man accumulating his selection successive slight variations will then pass the variability species state nature but shall unfortunately compelled treat this subject far too briefly can treated properly only giving long catalogues facts shall however enabled discuss what circumstances are most favourable variation the next chapter the struggle for existence amongst all organic beings throughout the world which inevitably follows from their high geometrical powers increase will treated this the doctrine malthus applied the whole animal and vegetable kingdoms many more individuals each species are born than can possibly survive and consequently there frequently recurring struggle for existence follows that any being vary however slightly any manner profitable itself under the complex and sometimes varying conditions life will have better chance surviving and thus naturally selected from the strong principle inheritance any selected variety will tend propagate its new and modified form this fundamental subject natural selection will treated some length the fourth chapter and shall then see how natural selection almost inevitably causes much extinction the less improved forms life and induces what have called divergence character the next chapter shall discuss the complex and little known laws variation and correlation growth the four succeeding chapters the most apparent and gravest difficulties the theory will given namely first the difficulties transitions understanding how simple being simple organ can changed and perfected into highly developed being elaborately constructed organ secondly the subject instinct the mental powers animals thirdly hybridism the infertility species and the fertility varieties when intercrossed and fourthly the imperfection the geological record the next chapter shall consider the geological succession organic beings throughout time the eleventh and twelfth their geographical distribution throughout space the thirteenth their classification mutual affinities both when mature and embryonic condition the last chapter shall give brief recapitulation the whole work and few concluding remarks one ought feel surprise much remaining yet unexplained regard the origin species and varieties makes due allowance for our profound ignorance regard the mutual relations all the beings which live around who can explain why one species ranges widely and very numerous and why another allied species has narrow range and rare yet these relations are the highest importance for they determine the present welfare and believe the future success and modification every inhabitant this world still less know the mutual relations the innumerable inhabitants the world during the many past geological epochs its history although much remains obscure and will long remain obscure can entertain doubt after the most deliberate study and dispassionate judgment which capable that the view which most naturalists entertain and which formerly entertained namely that each species has been independently created erroneous fully convinced that species are not immutable but that those belonging what are called the same genera are lineal descendants some other and generally extinct species the same manner the acknowledged varieties any one species are the descendants that species furthermore convinced that natural selection has been the main but not exclusive means modification chapter variation under domestication causes variability effects habit correlation growth inheritance character domestic varieties difficulty distinguishing between varieties and species origin domestic varieties from one more species domestic pigeons their differences and origin principle selection anciently followed its effects methodical and unconscious selection unknown origin our domestic productions circumstances favourable man power selection when look the individuals the same variety sub variety our older cultivated plants and animals one the first points which strikes that they generally differ much more from each other than the individuals any one species variety state nature when reflect the vast diversity the plants and animals which have been cultivated and which have varied during all ages under the most different climates and treatment think are driven conclude that this greater variability simply due our domestic productions having been raised under conditions life not uniform and somewhat different from those which the parent species have been exposed under nature there also think some probability the view propounded andrew knight that this variability may partly connected with excess food seems pretty clear that organic beings must exposed during several generations the new conditions life cause any appreciable amount variation and that when the organisation has once begun vary generally continues vary for many generations case record variable being ceasing variable under cultivation our oldest cultivated plants such wheat still often yield new varieties our oldest domesticated animals are still capable rapid improvement modification has been disputed what period life the causes variability whatever they may generally act whether during the early late period development the embryo the instant conception geoffroy hilaire experiments show that unnatural treatment the embryo causes monstrosities and monstrosities cannot separated any clear line distinction from mere variations but strongly inclined suspect that the most frequent cause variability may attributed the male and female reproductive elements having been affected prior the act conception several reasons make believe this but the chief one the remarkable effect which confinement cultivation has the functions the reproductive system this system appearing far more susceptible than any other part the organisation the action any change the conditions life nothing more easy than tame animal and few things more difficult than get breed freely under confinement even the many cases when the male and female unite how many animals there are which will not breed though living long under not very close confinement their native country this generally attributed vitiated instincts but how many cultivated plants display the utmost vigour and yet rarely never seed some few such cases has been found out that very trifling changes such little more less water some particular period growth will determine whether not the plant sets seed cannot here enter the copious details which have collected this curious subject but show how singular the laws are which determine the reproduction animals under confinement may just mention that carnivorous animals even from the tropics breed this country pretty freely under confinement with the exception the plantigrades bear family whereas carnivorous birds with the rarest exceptions hardly ever lay fertile eggs many exotic plants have pollen utterly worthless the same exact condition the most sterile hybrids when the one hand see domesticated animals and plants though often weak and sickly yet breeding quite freely under confinement and when the other hand see individuals though taken young from state nature perfectly tamed long lived and healthy which could give numerous instances yet having their reproductive system seriously affected unperceived causes fail acting need not surprised this system when does act under confinement acting not quite regularly and producing offspring not perfectly like their parents variable sterility has been said the bane horticulture but this view owe variability the same cause which produces sterility and variability the source all the choicest productions the garden may add that some organisms will breed most freely under the most unnatural conditions for instance the rabbit and ferret kept hutches showing that their reproductive system has not been thus affected will some animals and plants withstand domestication cultivation and vary very slightly perhaps hardly more than state nature long list could easily given sporting plants this term gardeners mean single bud offset which suddenly assumes new and sometimes very different character from that the rest the plant such buds can propagated grafting etc and sometimes seed these sports are extremely rare under nature but far from rare under cultivation and this case see that the treatment the parent has affected bud offset and not the ovules pollen but the opinion most physiologists that there essential difference between bud and ovule their earliest stages formation that fact sports support view that variability may largely attributed the ovules pollen both having been affected the treatment the parent prior the act conception these cases anyhow show that variation not necessarily connected some authors have supposed with the act generation seedlings from the same fruit and the young the same litter sometimes differ considerably from each other though both the young and the parents müller has remarked have apparently been exposed exactly the same conditions life and this shows how unimportant the direct effects the conditions life are comparison with the laws reproduction and growth and inheritance for had the action the conditions been direct any the young had varied all would probably have varied the same manner judge how much the case any variation should attribute the direct action heat moisture light food etc most difficult impression that with animals such agencies have produced very little direct effect though apparently more the case plants under this point view buckman recent experiments plants seem extremely valuable when all nearly all the individuals exposed certain conditions are affected the same way the change first appears directly due such conditions but some cases can shown that quite opposite conditions produce similar changes structure nevertheless some slight amount change may think attributed the direct action the conditions life some cases increased size from amount food colour from particular kinds food and from light and perhaps the thickness fur from climate habit also has decided influence the period flowering with plants when transported from one climate another animals has more marked effect for instance find the domestic duck that the bones the wing weigh less and the bones the leg more proportion the whole skeleton than the same bones the wild duck and presume that this change may safely attributed the domestic duck flying much less and walking more than its wild parent the great and inherited development the udders cows and goats countries where they are habitually milked comparison with the state these organs other countries another instance the effect use not single domestic animal can named which has not some country drooping ears and the view suggested some authors that the drooping due the disuse the muscles the ear from the animals not being much alarmed danger seems probable there are many laws regulating variation some few which can dimly seen and will hereafter briefly mentioned will here only allude what may called correlation growth any change the embryo larva will almost certainly entail changes the mature animal monstrosities the correlations between quite distinct parts are very curious and many instances are given isidore geoffroy hilaire great work this subject breeders believe that long limbs are almost always accompanied elongated head some instances correlation are quite whimsical thus cats with blue eyes are invariably deaf colour and constitutional peculiarities together which many remarkable cases could given amongst animals and plants from the facts collected heusinger appears that white sheep and pigs are differently affected from coloured individuals certain vegetable poisons hairless dogs have imperfect teeth long haired and coarse haired animals are apt have asserted long many horns pigeons with feathered feet have skin between their outer toes pigeons with short beaks have small feet and those with long beaks large feet hence man goes selecting and thus augmenting any peculiarity will almost certainly unconsciously modify other parts the structure owing the mysterious laws the correlation growth the result the various quite unknown dimly seen laws variation infinitely complex and diversified well worth while carefully study the several treatises published some our old cultivated plants the hyacinth potato even the dahlia etc and really surprising note the endless points structure and constitution which the varieties and sub varieties differ slightly from each other the whole organisation seems have become plastic and tends depart some small degree from that the parental type any variation which not inherited unimportant for but the number and diversity inheritable deviations structure both those slight and those considerable physiological importance endless prosper lucas treatise two large volumes the fullest and the best this subject breeder doubts how strong the tendency inheritance like produces like his fundamental belief doubts have been thrown this principle theoretical writers alone when deviation appears not unfrequently and see the father and child cannot tell whether may not due the same original cause acting both but when amongst individuals apparently exposed the same conditions any very rare deviation due some extraordinary combination circumstances appears the parent say once amongst several million individuals and reappears the child the mere doctrine chances almost compels attribute its reappearance inheritance every one must have heard cases albinism prickly skin hairy bodies etc appearing several members the same family strange and rare deviations structure are truly inherited less strange and commoner deviations may freely admitted inheritable perhaps the correct way viewing the whole subject would look the inheritance every character whatever the rule and non inheritance the anomaly the laws governing inheritance are quite unknown one can say why the same peculiarity different individuals the same species and individuals different species sometimes inherited and sometimes not why the child often reverts certain characters its grandfather grandmother other much more remote ancestor why peculiarity often transmitted from one sex both sexes one sex alone more commonly but not exclusively the like sex fact some little importance that peculiarities appearing the males our domestic breeds are often transmitted either exclusively much greater degree males alone much more important rule which think may trusted that whatever period life peculiarity first appears tends appear the offspring corresponding age though sometimes earlier many cases this could not otherwise thus the inherited peculiarities the horns cattle could appear only the offspring when nearly mature peculiarities the silkworm are known appear the corresponding caterpillar cocoon stage but hereditary diseases and some other facts make believe that the rule has wider extension and that when there apparent reason why peculiarity should appear any particular age yet that does tend appear the offspring the same period which first appeared the parent believe this rule the highest importance explaining the laws embryology these remarks are course confined the first appearance the peculiarity and not its primary cause which may have acted the ovules male element nearly the same manner the crossed offspring from short horned cow long horned bull the greater length horn though appearing late life clearly due the male element having alluded the subject reversion may here refer statement often made naturalists namely that our domestic varieties when run wild gradually but certainly revert character their aboriginal stocks hence has been argued that deductions can drawn from domestic races species state nature have vain endeavoured discover what decisive facts the above statement has often and boldly been made there would great difficulty proving its truth may safely conclude that very many the most strongly marked domestic varieties could not possibly live wild state many cases not know what the aboriginal stock was and could not tell whether not nearly perfect reversion had ensued would quite necessary order prevent the effects intercrossing that only single variety should turned loose its new home nevertheless our varieties certainly occasionally revert some their characters ancestral forms seems not improbable that could succeed naturalising were cultivate during many generations the several races for instance the cabbage very poor soil which case however some effect would have attributed the direct action the poor soil that they would large extent even wholly revert the wild aboriginal stock whether not the experiment would succeed not great importance for our line argument for the experiment itself the conditions life are changed could shown that our domestic varieties manifested strong tendency reversion that lose their acquired characters whilst kept under unchanged conditions and whilst kept considerable body that free intercrossing might check blending together any slight deviations structure such case grant that could deduce nothing from domestic varieties regard species but there not shadow evidence favour this view assert that could not breed our cart and race horses long and short horned cattle and poultry various breeds and esculent vegetables for almost infinite number generations would opposed all experience may add that when under nature the conditions life change variations and reversions character probably occur but natural selection will hereafter explained will determine how far the new characters thus arising shall preserved when look the hereditary varieties races our domestic animals and plants and compare them with species closely allied together generally perceive each domestic race already remarked less uniformity character than true species domestic races the same species also often have somewhat monstrous character which mean that although differing from each other and from the other species the same genus several trifling respects they often differ extreme degree some one part both when compared one with another and more especially when compared with all the species nature which they are nearest allied with these exceptions and with that the perfect fertility varieties when crossed subject hereafter discussed domestic races the same species differ from each other the same manner only most cases lesser degree than closely allied species the same genus state nature think this must admitted when find that there are hardly any domestic races either amongst animals plants which have not been ranked some competent judges mere varieties and other competent judges the descendants aboriginally distinct species any marked distinction existed between domestic races and species this source doubt could not perpetually recur has often been stated that domestic races not differ from each other characters generic value think could shown that this statement hardly correct but naturalists differ most widely determining what characters are generic value all such valuations being present empirical moreover the view the origin genera which shall presently give have right expect often meet with generic differences our domesticated productions when attempt estimate the amount structural difference between the domestic races the same species are soon involved doubt from not knowing whether they have descended from one several parent species this point could cleared would interesting for instance could shown that the greyhound bloodhound terrier spaniel and bull dog which all know propagate their kind truly were the offspring any single species then such facts would have great weight making doubt about the immutability the many very closely allied and natural species for instance the many foxes inhabiting different quarters the world not believe shall presently see that all our dogs have descended from any one wild species but the case some other domestic races there presumptive even strong evidence favour this view has often been assumed that man has chosen for domestication animals and plants having extraordinary inherent tendency vary and likewise withstand diverse climates not dispute that these capacities have added largely the value most our domesticated productions but how could savage possibly know when first tamed animal whether would vary succeeding generations and whether would endure other climates has the little variability the ass guinea fowl the small power endurance warmth the rein deer cold the common camel prevented their domestication cannot doubt that other animals and plants equal number our domesticated productions and belonging equally diverse classes and countries were taken from state nature and could made breed for equal number generations under domestication they would vary average largely the parent species our existing domesticated productions have varied the case most our anciently domesticated animals and plants not think possible come any definite conclusion whether they have descended from one several species the argument mainly relied those who believe the multiple origin our domestic animals that find the most ancient records more especially the monuments egypt much diversity the breeds and that some the breeds closely resemble perhaps are identical with those still existing even this latter fact were found more strictly and generally true than seems the case what does show but that some our breeds originated there four five thousand years ago but horner researches have rendered some degree probable that man sufficiently civilized have manufactured pottery existed the valley the nile thirteen fourteen thousand years ago and who will pretend say how long before these ancient periods savages like those tierra del fuego australia who possess semi domestic dog may not have existed egypt the whole subject must think remain vague nevertheless may without here entering any details state that from geographical and other considerations think highly probable that our domestic dogs have descended from several wild species regard sheep and goats can form opinion should think from facts communicated blyth the habits voice and constitution etc the humped indian cattle that these had descended from different aboriginal stock from our european cattle and several competent judges believe that these latter have had more than one wild parent with respect horses from reasons which cannot give here doubtfully inclined believe opposition several authors that all the races have descended from one wild stock blyth whose opinion from his large and varied stores knowledge should value more than that almost any one thinks that all the breeds poultry have proceeded from the common wild indian fowl gallus bankiva regard ducks and rabbits the breeds which differ considerably from each other structure not doubt that they all have descended from the common wild duck and rabbit the doctrine the origin our several domestic races from several aboriginal stocks has been carried absurd extreme some authors they believe that every race which breeds true let the distinctive characters ever slight has had its wild prototype this rate there must have existed least score species wild cattle many sheep and several goats europe alone and several even within great britain one author believes that there formerly existed great britain eleven wild species sheep peculiar when bear mind that britain has now hardly one peculiar mammal and france but few distinct from those germany and conversely and with hungary spain etc but that each these kingdoms possesses several peculiar breeds cattle sheep etc must admit that many domestic breeds have originated europe for whence could they have been derived these several countries not possess number peculiar species distinct parent stocks india even the case the domestic dogs the whole world which fully admit have probably descended from several wild species cannot doubt that there has been immense amount inherited variation who can believe that animals closely resembling the italian greyhound the bloodhound the bull dog blenheim spaniel etc unlike all wild canidæ ever existed freely state nature has often been loosely said that all our races dogs have been produced the crossing few aboriginal species but crossing can get only forms some degree intermediate between their parents and account for our several domestic races this process must admit the former existence the most extreme forms the italian greyhound bloodhound bull dog etc the wild state moreover the possibility making distinct races crossing has been greatly exaggerated there can doubt that race may modified occasional crosses aided the careful selection those individual mongrels which present any desired character but that race could obtained nearly intermediate between two extremely different races species can hardly believe sir sebright expressly experimentised for this object and failed the offspring from the first cross between two pure breeds tolerably and sometimes have found with pigeons extremely uniform and everything seems simple enough but when these mongrels are crossed one with another for several generations hardly two them will alike and then the extreme difficulty rather utter hopelessness the task becomes apparent certainly breed intermediate between two very distinct breeds could not got without extreme care and long continued selection nor can find single case record permanent race having been thus formed the breeds the domestic pigeon believing that always best study some special group have after deliberation taken domestic pigeons have kept every breed which could purchase obtain and have been most kindly favoured with skins from several quarters the world more especially the honourable elliot from india and the honourable murray from persia many treatises different languages have been published pigeons and some them are very important being considerable antiquity have associated with several eminent fanciers and have been permitted join two the london pigeon clubs the diversity the breeds something astonishing compare the english carrier and the short faced tumbler and see the wonderful difference their beaks entailing corresponding differences their skulls the carrier more especially the male bird also remarkable from the wonderful development the carunculated skin about the head and this accompanied greatly elongated eyelids very large external orifices the nostrils and wide gape mouth the short faced tumbler has beak outline almost like that finch and the common tumbler has the singular and strictly inherited habit flying great height compact flock and tumbling the air head over heels the runt bird great size with long massive beak and large feet some the sub breeds runts have very long necks others very long wings and tails others singularly short tails the barb allied the carrier but instead very long beak has very short and very broad one the pouter has much elongated body wings and legs and its enormously developed crop which glories inflating may well excite astonishment and even laughter the turbit has very short and conical beak with line reversed feathers down the breast and has the habit continually expanding slightly the upper part the oesophagus the jacobin has the feathers much reversed along the back the neck that they form hood and has proportionally its size much elongated wing and tail feathers the trumpeter and laugher their names express utter very different coo from the other breeds the fantail has thirty even forty tail feathers instead twelve fourteen the normal number all members the great pigeon family and these feathers are kept expanded and are carried erect that good birds the head and tail touch the oil gland quite aborted several other less distinct breeds might have been specified the skeletons the several breeds the development the bones the face length and breadth and curvature differs enormously the shape well the breadth and length the ramus the lower jaw varies highly remarkable manner the number the caudal and sacral vertebræ vary does the number the ribs together with their relative breadth and the presence processes the size and shape the apertures the sternum are highly variable the degree divergence and relative size the two arms the furcula the proportional width the gape mouth the proportional length the eyelids the orifice the nostrils the tongue not always strict correlation with the length beak the size the crop and the upper part the oesophagus the development and abortion the oil gland the number the primary wing and caudal feathers the relative length wing and tail each other and the body the relative length leg and the feet the number scutellæ the toes the development skin between the toes are all points structure which are variable the period which the perfect plumage acquired varies does the state the down with which the nestling birds are clothed when hatched the shape and size the eggs vary the manner flight differs remarkably does some breeds the voice and disposition lastly certain breeds the males and females have come differ slight degree from each other altogether least score pigeons might chosen which shown ornithologist and were told that they were wild birds would certainly think ranked him well defined species moreover not believe that any ornithologist would place touch the oil gland quite aborted several other less distinct breeds might have been specified the english carrier the short faced tumbler the runt the barb pouter and fantail the same genus more especially each these breeds several truly inherited sub breeds species might have called them could shown him great the differences are between the breeds pigeons fully convinced that the common opinion naturalists correct namely that all have descended from the rock pigeon columba livia including under this term several geographical races sub species which differ from each other the most trifling respects several the reasons which have led this belief are some degree applicable other cases will here briefly give them the several breeds are not varieties and have not proceeded from the rock pigeon they must have descended from least seven eight aboriginal stocks for impossible make the present domestic breeds the crossing any lesser number how for instance could pouter produced crossing two breeds unless one the parent stocks possessed the characteristic enormous crop the supposed aboriginal stocks must all have been rock pigeons that not breeding willingly perching trees but besides livia with its geographical sub species only two three other species rock pigeons are known and these have not any the characters the domestic breeds hence the supposed aboriginal stocks must either still exist the countries where they were originally domesticated and yet unknown ornithologists and this considering their size habits and remarkable characters seems very improbable they must have become extinct the wild state but birds breeding precipices and good fliers are unlikely exterminated and the common rock pigeon which has the same habits with the domestic breeds has not been exterminated even several the smaller british islets the shores the mediterranean hence the supposed extermination many species having similar habits with the rock pigeon seems very rash assumption moreover the several above named domesticated breeds have been transported all parts the world and therefore some them must have been carried back again into their native country but not one has ever become wild feral though the dovecot pigeon which the rock pigeon very slightly altered state has become feral several places again all recent experience shows that most difficult get any wild animal breed freely under domestication yet the hypothesis the multiple origin our pigeons must assumed that least seven eight species were thoroughly domesticated ancient times half civilized man quite prolific under confinement argument seems great weight and applicable several other cases that the above specified breeds though agreeing generally constitution habits voice colouring and most parts their structure with the wild rock pigeon yet are certainly highly abnormal other parts their structure may look vain throughout the whole great family columbidæ for beak like that the english carrier that the short faced tumbler barb for reversed feathers like those the jacobin for crop like that the pouter for tail feathers like those the fantail hence must assumed not only that half civilized man succeeded thoroughly domesticating several species but that intentionally chance picked out extraordinarily abnormal species and further that these very species have since all become extinct unknown many strange contingencies seem improbable the highest degree some facts regard the colouring pigeons well deserve consideration the rock pigeon slaty blue and has white rump the indian sub species intermedia strickland having bluish the tail has terminal dark bar with the bases the outer feathers externally edged with white the wings have two black bars some semi domestic breeds and some apparently truly wild breeds have besides the two black bars the wings chequered with black these several marks not occur together any other species the whole family now every one the domestic breeds taking thoroughly well bred birds all the above marks even the white edging the outer tail feathers sometimes concur perfectly developed moreover when two birds belonging two distinct breeds are crossed neither which blue has any the above specified marks the mongrel offspring are very apt suddenly acquire these characters for instance crossed some uniformly white fantails with some uniformly black barbs and they produced mottled brown and black birds these again crossed together and one grandchild the pure white fantail and pure black barb was beautiful blue colour with the white rump double black wing bar and barred and white edged tail feathers any wild rock pigeon can understand these facts the well known principle reversion ancestral characters all the domestic breeds have descended from the rock pigeon but deny this must make one the two following highly improbable suppositions either firstly that all the several imagined aboriginal stocks were coloured and marked like the rock pigeon although other existing species thus coloured and marked that each separate breed there might tendency revert the very same colours and markings secondly that each breed even the purest has within dozen most within score generations been crossed the rock pigeon say within dozen twenty generations for know fact countenancing the belief that the child ever reverts some one ancestor removed greater number generations breed which has been crossed only once with some distinct breed the tendency reversion any character derived from such cross will naturally become less and less each succeeding generation there will less the foreign blood but when there has been cross with distinct breed and there tendency both parents revert character which has been lost during some former generation this tendency for all that can see the contrary may transmitted undiminished for indefinite number generations these two distinct cases are often confounded treatises inheritance lastly the hybrids mongrels from between all the domestic breeds pigeons are perfectly fertile can state this from own observations purposely made the most distinct breeds now difficult perhaps impossible bring forward one case the hybrid offspring two animals clearly distinct being themselves perfectly fertile some authors believe that long continued domestication eliminates this strong tendency sterility from the history the dog think there some probability this hypothesis applied species closely related together though unsupported single experiment but extend the hypothesis far suppose that species aboriginally distinct carriers tumblers pouters and fantails now are should yield offspring perfectly fertile inter seems rash the extreme from these several reasons namely the improbability man having formerly got seven eight supposed species pigeons breed freely under domestication these supposed species being quite unknown wild state and their becoming nowhere feral these species having very abnormal characters certain respects compared with all other columbidæ though like most other respects the rock pigeon the blue colour and various marks occasionally appearing all the breeds both when kept pure and when crossed the mongrel offspring being perfectly fertile from these several reasons taken together can feel doubt that all our domestic breeds have descended from the columba livia with its geographical sub species favour this view may add firstly that livia the rock pigeon has been found capable domestication europe and india and that agrees habits and great number points structure with all the domestic breeds secondly although english carrier short faced tumbler differs immensely certain characters from the rock pigeon yet comparing the several sub breeds these breeds more especially those brought from distant countries can make almost perfect series between the extremes structure thirdly those characters which are mainly distinctive each breed for instance the wattle and length beak the carrier the shortness that the tumbler and the number tail feathers the fantail are each breed eminently variable and the explanation this fact will obvious when come treat selection fourthly pigeons have been watched and tended with the utmost care and loved many people they have been domesticated for thousands years several quarters the world the earliest known record pigeons the fifth aegyptian dynasty about 3000 was pointed out professor lepsius but birch informs that pigeons are given bill fare the previous dynasty the time the romans hear from pliny immense prices were given for pigeons nay they are come this pass that they can reckon their pedigree and race pigeons were much valued akber khan india about the year 1600 never less than 000 pigeons were taken with the court the monarchs iran and turan sent him some very rare birds and continues the courtly historian his majesty crossing the breeds which method was never practised before has improved them astonishingly about this same period the dutch were eager about pigeons were the old romans the paramount importance these considerations explaining the immense amount variation which pigeons have undergone will obvious when treat selection shall then also see how that the breeds often have somewhat monstrous character also most favourable circumstance for the production distinct breeds that male and female pigeons can easily mated for life and thus different breeds can kept together the same aviary have discussed the probable origin domestic pigeons some yet quite insufficient length because when first kept pigeons and watched the several kinds knowing well how true they bred felt fully much difficulty believing that they could ever have descended from common parent any naturalist could coming similar conclusion regard the many species finches other large groups birds nature one circumstance has struck much namely that all the breeders the various domestic animals and the cultivators plants with whom have ever conversed whose treatises have read are firmly convinced that the several breeds which each has attended are descended from many aboriginally distinct species ask have asked celebrated raiser hereford cattle whether his cattle might not have descended from long horns and will laugh you scorn have never met pigeon poultry duck rabbit fancier who was not fully convinced that each main breed was descended from distinct species van mons his treatise pears and apples shows how utterly disbelieves that the several sorts for instance ribston pippin codlin apple could ever have proceeded from the seeds the same tree innumerable other examples could given the explanation think simple from long continued study they are strongly impressed with the differences between the several races and though they well know that each race varies slightly for they win their prizes selecting such slight differences yet they ignore all general arguments and refuse sum their minds slight differences accumulated during many successive generations may not those naturalists who knowing far less the laws inheritance than does the breeder and knowing more than does the intermediate links the long lines descent yet admit that many our domestic races have descended from the same parents may they not learn lesson caution when they deride the idea species state nature being lineal descendants other species selection let now briefly consider the steps which domestic races have been produced either from one from several allied species some little effect may perhaps attributed the direct action the external conditions life and some little habit but would bold man who would account such agencies for the differences dray and race horse greyhound and bloodhound carrier and tumbler pigeon one the most remarkable features our domesticated races that see them adaptation not indeed the animal plant own good but man use fancy some variations useful him have probably arisen suddenly one step many botanists for instance believe that the fuller teazle with its hooks which cannot rivalled any mechanical contrivance only variety the wild dipsacus and this amount change may have suddenly arisen seedling has probably been with the turnspit dog and this known have been the case with the ancon sheep but when compare the dray horse and race horse the dromedary and camel the various breeds sheep fitted either for cultivated land mountain pasture with the wool one breed good for one purpose and that another breed for another purpose when compare the many breeds dogs each good for man very different ways when compare the game cock pertinacious battle with other breeds little quarrelsome with everlasting layers which never desire sit and with the bantam small and elegant when compare the host agricultural culinary orchard and flower garden races plants most useful man different seasons and for different purposes beautiful his eyes must think look further than mere variability cannot suppose that all the breeds were suddenly produced perfect and useful now see them indeed several cases know that this has not been their history the key man power accumulative selection nature gives successive variations man adds them certain directions useful him this sense may said make for himself useful breeds the great power this principle selection not hypothetical certain that several our eminent breeders have even within single lifetime modified large extent some breeds cattle and sheep order fully realise what they have done almost necessary read several the many treatises devoted this subject and inspect the animals breeders habitually speak animal organisation something quite plastic which they can model almost they please had space could quote numerous passages this effect from highly competent authorities youatt who was probably better acquainted with the works agriculturalists than almost any other individual and who was himself very good judge animal speaks the principle selection that which enables the agriculturist not only modify the character his flock but change altogether the magician wand means which may summon into life whatever form and mould pleases lord somerville speaking what breeders have done for sheep says would seem they had chalked out upon wall form perfect itself and then had given existence that most skilful breeder sir john sebright used say with respect pigeons that would produce any given feather three years but would take him six years obtain head and beak saxony the importance the principle selection regard merino sheep fully recognised that men follow trade the sheep are placed table and are studied like picture connoisseur this done three times intervals months and the sheep are each time marked and classed that the very best may ultimately selected for breeding what english breeders have actually effected proved the enormous prices given for animals with good pedigree and these have now been exported almost every quarter the world the improvement means generally due crossing different breeds all the best breeders are strongly opposed this practice except sometimes amongst closely allied sub breeds and when cross has been made the closest selection far more indispensable even than ordinary cases selection consisted merely separating some very distinct variety and breeding from the principle would obvious hardly worth notice but its importance consists the great effect produced the accumulation one direction during successive generations differences absolutely inappreciable uneducated eye differences which for one have vainly attempted appreciate not one man thousand has accuracy eye and judgment sufficient become eminent breeder gifted with these qualities and studies his subject for years and devotes his lifetime with indomitable perseverance will succeed and may make great improvements wants any these qualities will assuredly fail few would readily believe the natural capacity and years practice requisite become even skilful pigeon fancier the same principles are followed horticulturists but the variations are here often more abrupt one supposes that our choicest productions have been produced single variation from the aboriginal stock have proofs that this not some cases which exact records have been kept thus give very trifling instance the steadily increasing size the common gooseberry may quoted see astonishing improvement many florists flowers when the flowers the present day are compared with drawings made only twenty thirty years ago when race plants once pretty well established the seed raisers not pick out the best plants but merely over their seed beds and pull the rogues they call the plants that deviate from the proper standard with animals this kind selection fact also followed for hardly any one careless allow his worst animals breed regard plants there another means observing the accumulated effects selection namely comparing the diversity flowers the different varieties the same species the flower garden the diversity leaves pods tubers whatever part valued the kitchen garden comparison with the flowers the same varieties and the diversity fruit the same species the orchard comparison with the leaves and flowers the same set varieties see how different the leaves the cabbage are and how extremely alike the flowers how unlike the flowers the heartsease are and how alike the leaves how much the fruit the different kinds gooseberries differ size colour shape and hairiness and yet the flowers present very slight differences not that the varieties which differ largely some one point not differ all other points this hardly ever perhaps never the case the laws correlation growth the importance which should never overlooked will ensure some differences but general rule cannot doubt that the continued selection slight variations either the leaves the flowers the fruit will produce races differing from each other chiefly these characters may objected that the principle selection has been reduced methodical practice for scarcely more than three quarters century has certainly been more attended late years and many treatises have been published the subject and the result may add has been corresponding degree rapid and important but very far from true that the principle modern discovery could give several references the full acknowledgment the importance the principle works high antiquity rude and barbarous periods english history choice animals were often imported and laws were passed prevent their exportation the destruction horses under certain size was ordered and this may compared the roguing plants nurserymen the principle selection find distinctly given ancient chinese encyclopædia explicit rules are laid down some the roman classical writers from passages genesis clear that the colour domestic animals was that early period attended savages now sometimes cross their dogs with wild canine animals improve the breed and they formerly did attested passages pliny the savages south africa match their draught cattle colour some the esquimaux their teams dogs livingstone shows how much good domestic breeds are valued the negroes the interior africa who have not associated with europeans some these facts not show actual selection but they show that the breeding domestic animals was carefully attended ancient times and now attended the lowest savages would indeed have been strange fact had attention not been paid breeding for the inheritance good and bad qualities obvious the present time eminent breeders try methodical selection with distinct object view make new strain sub breed superior anything existing the country but for our purpose kind selection which may called unconscious and which results from every one trying possess and breed from the best individual animals more important thus man who intends keeping pointers naturally tries get good dogs can and afterwards breeds from his own best dogs but has wish expectation permanently altering the breed nevertheless cannot doubt that this process continued during centuries would improve and modify any breed the same way bakewell collins etc this very same process only carried more methodically did greatly modify even during their own lifetimes the forms and qualities their cattle slow and insensible changes this kind could never recognised unless actual measurements careful drawings the breeds question had been made long ago which might serve for comparison some cases however unchanged but little changed individuals the same breed may found less civilised districts where the breed has been less improved there reason believe that king charles spaniel has been unconsciously modified large extent since the time that monarch some highly competent authorities are convinced that the setter directly derived from the spaniel and has probably been slowly altered from known that the english pointer has been greatly changed within the last century and this case the change has believed been chiefly effected crosses with the fox hound but what concerns that the change has been effected unconsciously and gradually and yet effectually that though the old spanish pointer certainly came from spain borrow has not seen informed him any native dog spain like our pointer similar process selection and careful training the whole body english racehorses have come surpass fleetness and size the parent arab stock that the latter the regulations for the goodwood races are favoured the weights they carry lord spencer and others have shown how the cattle england have increased weight and early maturity compared with the stock formerly kept this country comparing the accounts given old pigeon treatises carriers and tumblers with these breeds now existing britain india and persia can think clearly trace the stages through which they have insensibly passed and come differ greatly from the rock pigeon youatt gives excellent illustration the effects course selection which may considered unconsciously followed far that the breeders could never have expected even have wished have produced the result which ensued namely the production two distinct strains the two flocks leicester sheep kept buckley and burgess youatt remarks have been purely bred from the original stock bakewell for upwards fifty years there not suspicion existing the mind any one all acquainted with the subject that the owner either them has deviated any one instance from the pure blood bakewell flock and yet the difference between the sheep possessed these two gentlemen great that they have the appearance being quite different varieties there exist savages barbarous never think the inherited character the offspring their domestic animals yet any one animal particularly useful them for any special purpose would carefully preserved during famines and other accidents which savages are liable and such choice animals would thus generally leave more offspring than the inferior ones that this case there would kind unconscious selection going see the value set animals even the barbarians tierra del fuego their killing and devouring their old women times dearth less value than their dogs plants the same gradual process improvement through the occasional preservation the best individuals whether not sufficiently distinct ranked their first appearance distinct varieties and whether not two more species races have become blended together crossing may plainly recognised the increased size and beauty which now see the varieties the heartsease rose pelargonium dahlia and other plants when compared with the older varieties with their parent stocks one would ever expect get first rate heartsease dahlia from the seed wild plant one would expect raise first rate melting pear from the seed wild pear though might succeed from poor seedling growing wild had come from garden stock the pear though cultivated classical times appears from pliny description have been fruit very inferior quality have seen great surprise expressed horticultural works the wonderful skill gardeners having produced such splendid results from such poor materials but the art cannot doubt has been simple and far the final result concerned has been followed almost unconsciously has consisted always cultivating the best known variety sowing its seeds and when slightly better variety has chanced appear selecting and onwards but the gardeners the classical period who cultivated the best pear they could procure never thought what splendid fruit should eat though owe our excellent fruit some small degree their having naturally chosen and preserved the best varieties they could anywhere find large amount change our cultivated plants thus slowly and unconsciously accumulated explains believe the well known fact that vast number cases cannot recognise and therefore not know the wild parent stocks the plants which have been longest cultivated our flower and kitchen gardens has taken centuries thousands years improve modify most our plants their present standard usefulness man can understand how that neither australia the cape good hope nor any other region inhabited quite uncivilised man has afforded single plant worth culture not that these countries rich species not strange chance possess the aboriginal stocks any useful plants but that the native plants have not been improved continued selection standard perfection comparable with that given the plants countries anciently civilised regard the domestic animals kept uncivilised man should not overlooked that they almost always have struggle for their own food least during certain seasons and two countries very differently circumstanced individuals the same species having slightly different constitutions structure would often succeed better the one country than the other and thus process natural selection will hereafter more fully explained two sub breeds might formed this perhaps partly explains what has been remarked some authors namely that the varieties kept savages have more the character species than the varieties kept civilised countries the view here given the all important part which selection man has played becomes once obvious how that our domestic races show adaptation their structure their habits man wants fancies can think further understand the frequently abnormal character our domestic races and likewise their differences being great external characters and relatively slight internal parts organs man can hardly select only with much difficulty any deviation structure excepting such externally visible and indeed rarely cares for what internal can never act selection excepting variations which are first given him some slight degree nature man would ever try make fantail till saw pigeon with tail developed some slight degree unusual manner pouter till saw pigeon with crop somewhat unusual size and the more abnormal unusual any character was when first appeared the more likely would catch his attention but use such expression trying make fantail have doubt most cases utterly incorrect the man who first selected pigeon with slightly larger tail never dreamed what the descendants that pigeon would become through long continued partly unconscious and partly methodical selection perhaps the parent bird all fantails had only fourteen tail feathers somewhat expanded like the present java fantail like individuals other and distinct breeds which many seventeen tail feathers have been counted perhaps the first pouter pigeon did not inflate its crop much more than the turbit now does the upper part its oesophagus habit which disregarded all fanciers not one the points the breed nor let thought that some great deviation structure would necessary catch the fancier eye perceives extremely small differences and human nature value any novelty however slight one own possession nor must the value which would formerly set any slight differences the individuals the same species judged the value which would now set them after several breeds have once fairly been established many slight differences might and indeed now arise amongst pigeons which are rejected faults deviations from the standard perfection each breed the common goose has not given rise any marked varieties hence the thoulouse and the common breed which differ only colour that most fleeting characters have lately been exhibited distinct our poultry shows think these views further explain what has sometimes been noticed namely that know nothing about the origin history any our domestic breeds but fact breed like dialect language can hardly said have had definite origin man preserves and breeds from individual with some slight deviation structure takes more care than usual matching his best animals and thus improves them and the improved individuals slowly spread the immediate neighbourhood but yet they will hardly have distinct name and from being only slightly valued their history will disregarded when further improved the same slow and gradual process they will spread more widely and will get recognised something distinct and valuable and will then probably first receive provincial name semi civilised countries with little free communication the spreading and knowledge any new sub breed will slow process soon the points value the new sub breed are once fully acknowledged the principle have called unconscious selection will always tend perhaps more one period than another the breed rises falls fashion perhaps more one district than another according the state civilisation the inhabitants slowly add the characteristic features the breed whatever they may but the chance will infinitely small any record having been preserved such slow varying and insensible changes must now say few words the circumstances favourable the reverse man power selection high degree variability obviously favourable freely giving the materials for selection work not that mere individual differences are not amply sufficient with extreme care allow the accumulation large amount modification almost any desired direction but variations manifestly useful pleasing man appear only occasionally the chance their appearance will much increased large number individuals being kept and hence this comes the highest importance success this principle marshall has remarked with respect the sheep parts yorkshire that they generally belong poor people and are mostly small lots they never can improved the other hand nurserymen from raising large stocks the same plants are generally far more successful than amateurs getting new and valuable varieties the keeping large number individuals species any country requires that the species should placed under favourable conditions life breed freely that country when the individuals any species are scanty all the individuals whatever their quality may will generally allowed breed and this will effectually prevent selection but probably the most important point all that the animal plant should highly useful man much valued him that the closest attention should paid even the slightest deviation the qualities structure each individual unless such attention paid nothing can effected have seen gravely remarked that was most fortunate that the strawberry began vary just when gardeners began attend closely this plant doubt the strawberry had always varied since was cultivated but the slight varieties had been neglected soon however gardeners picked out individual plants with slightly larger earlier better fruit and raised seedlings from them and again picked out the best seedlings and bred from them then there appeared aided some crossing with distinct species those many admirable varieties the strawberry which have been raised during the last thirty forty years the case animals with separate sexes facility preventing crosses important element success the formation new races least country which already stocked with other races this respect enclosure the land plays part wandering savages the inhabitants open plains rarely possess more than one breed the same species pigeons can mated for life and this great convenience the fancier for thus many races may kept true though mingled the same aviary and this circumstance must have largely favoured the improvement and formation new breeds pigeons may add can propagated great numbers and very quick rate and inferior birds may freely rejected when killed they serve for food the other hand cats from their nocturnal rambling habits cannot matched and although much valued women and children hardly ever see distinct breed kept such breeds sometimes see are almost always imported from some other country often from islands although not doubt that some domestic animals vary less than others yet the rarity absence distinct breeds the cat the donkey peacock goose etc may attributed main part selection not having been brought into play cats from the difficulty pairing them donkeys from only few being kept poor people and little attention paid their breeding peacocks from not being very easily reared and large stock not kept geese from being valuable only for two purposes food and feathers and more especially from pleasure having been felt the display distinct breeds sum the origin our domestic races animals and plants believe that the conditions life from their action the reproductive system are far the highest importance causing variability not believe that variability inherent and necessary contingency under all circumstances with all organic beings some authors have thought the effects variability are modified various degrees inheritance and reversion variability governed many unknown laws more especially that correlation growth something may attributed the direct action the conditions life something must attributed use and disuse the final result thus rendered infinitely complex some cases not doubt that the intercrossing species aboriginally distinct has played important part the origin our domestic productions when any country several domestic breeds have once been established their occasional intercrossing with the aid selection has doubt largely aided the formation new sub breeds but the importance the crossing varieties has believe been greatly exaggerated both regard animals and those plants which are propagated seed plants which are temporarily propagated cuttings buds etc the importance the crossing both distinct species and varieties immense for the cultivator here quite disregards the extreme variability both hybrids and mongrels and the frequent sterility hybrids but the cases plants not propagated seed are little importance for their endurance only temporary over all these causes change convinced that the accumulative action selection whether applied methodically and more quickly unconsciously and more slowly but more efficiently far the predominant power chapter variation under nature variability individual differences doubtful species wide ranging much diffused and common species vary most species the larger genera any country vary more than the species the smaller genera many the species the larger genera resemble varieties being very closely but unequally related each other and having restricted ranges before applying the principles arrived the last chapter organic beings state nature must briefly discuss whether these latter are subject any variation treat this subject all properly long catalogue dry facts should given but these shall reserve for future work nor shall here discuss the various definitions which have been given the term species one definition has yet satisfied all naturalists yet every naturalist knows vaguely what means when speaks species generally the term includes the unknown element distinct act creation the term variety almost equally difficult define but here community descent almost universally implied though can rarely proved have also what are called monstrosities but they graduate into varieties monstrosity presume meant some considerable deviation structure one part either injurious not useful the species and not generally propagated some authors use the term variation technical sense implying modification directly due the physical conditions life and variations this sense are supposed not inherited but who can say that the dwarfed condition shells the brackish waters the baltic dwarfed plants alpine summits the thicker fur animal from far northwards would not some cases inherited for least some few generations and this case presume that the form would called variety again have many slight differences which may called individual differences such are known frequently appear the offspring from the same parents which may presumed have thus arisen from being frequently observed the individuals the same species inhabiting the same confined locality one supposes that all the individuals the same species are cast the very same mould these individual differences are highly important for they afford materials for natural selection accumulate the same manner man can accumulate any given direction individual differences his domesticated productions these individual differences generally affect what naturalists consider unimportant parts but could show long catalogue facts that parts which must called important whether viewed under physiological classificatory point view sometimes vary the individuals the same species convinced that the most experienced naturalist would surprised the number the cases variability even important parts structure which could collect good authority have collected during course years should remembered that systematists are far from pleased finding variability important characters and that there are not many men who will laboriously examine internal and important organs and compare them many specimens the same species should never have expected that the branching the main nerves close the great central ganglion insect would have been variable the same species should have expected that changes this nature could have been effected only slow degrees yet quite recently lubbock has shown degree variability these main nerves coccus which may almost compared the irregular branching the stem tree this philosophical naturalist may add has also quite recently shown that the muscles the larvæ certain insects are very far from uniform authors sometimes argue circle when they state that important organs never vary for these same authors practically rank that character important some few naturalists have honestly confessed which does not vary and under this point view instance important part varying will ever found but under any other point view many instances assuredly can given there one point connected with individual differences which seems extremely perplexing refer those genera which have sometimes been called protean polymorphic which the species present inordinate amount variation and hardly two naturalists can agree which forms rank species and which varieties may instance rubus rosa and hieracium amongst plants several genera insects and several genera brachiopod shells most polymorphic genera some the species have fixed and definite characters genera which are polymorphic one country seem with some few exceptions polymorphic other countries and likewise judging from brachiopod shells former periods time these facts seem very perplexing for they seem show that this kind variability independent the conditions life inclined suspect that see these polymorphic genera variations points structure which are service disservice the species and which consequently have not been seized and rendered definite natural selection hereafter will explained those forms which possess some considerable degree the character species but which are closely similar some other forms are closely linked them intermediate gradations that naturalists not like rank them distinct species are several respects the most important for have every reason believe that many these doubtful and closely allied forms have permanently retained their characters their own country for long time for long far know have good and true species practically when naturalist can unite two forms together others having intermediate characters treats the one variety the other ranking the most common but sometimes the one first described the species and the other the variety but cases great difficulty which will not here enumerate sometimes occur deciding whether not rank one form variety another even when they are closely connected intermediate links nor will the commonly assumed hybrid nature the intermediate links always remove the difficulty very many cases however one form ranked variety another not because the intermediate links have actually been found but because analogy leads the observer suppose either that they now somewhere exist may formerly have existed and here wide door for the entry doubt and conjecture opened hence determining whether form should ranked species variety the opinion naturalists having sound judgment and wide experience seems the only guide follow must however many cases decide majority naturalists for few well marked and well known varieties can named which have not been ranked species least some competent judges that varieties this doubtful nature are far from uncommon cannot disputed compare the several floras great britain france the united states drawn different botanists and see what surprising number forms have been ranked one botanist good species and another mere varieties watson whom lie under deep obligation for assistance all kinds has marked for 182 british plants which are generally considered varieties but which have all been ranked botanists species and making this list has omitted many trifling varieties but which nevertheless have been ranked some botanists species and has entirely omitted several highly polymorphic genera under genera including the most polymorphic forms babington gives 251 species whereas bentham gives only 112 difference 139 doubtful forms amongst animals which unite for each birth and which are highly locomotive doubtful forms ranked one zoologist species and another variety can rarely found within the same country but are common separated areas how many those birds and insects north america and europe which differ very slightly from each other have been ranked one eminent naturalist undoubted species and another varieties they are often called geographical races many years ago when comparing and seeing others compare the birds from the separate islands the galapagos archipelago both one with another and with those from the american mainland was much struck how entirely vague and arbitrary the distinction between species and varieties the islets the little madeira group there are many insects which are characterized varieties wollaston admirable work but which cannot doubted would ranked distinct species many entomologists even ireland has few animals now generally regarded varieties but which have been ranked species some zoologists several most experienced ornithologists consider our british red grouse only strongly marked race norwegian species whereas the greater number rank undoubted species peculiar great britain wide distance between the homes two doubtful forms leads many naturalists rank both distinct species but what distance has been well asked will suffice that between america and europe ample will that between the continent and the azores madeira the canaries ireland sufficient must admitted that many forms considered highly competent judges varieties have perfectly the character species that they are ranked other highly competent judges good and true species but discuss whether they are rightly called species varieties before any definition these terms has been generally accepted vainly beat the air many the cases strongly marked varieties doubtful species well deserve consideration for several interesting lines argument from geographical distribution analogical variation hybridism etc have been brought bear the attempt determine their rank will here give only single instance the well known one the primrose and cowslip primula veris and elatior these plants differ considerably appearance they have different flavour and emit different odour they flower slightly different periods they grow somewhat different stations they ascend mountains different heights they have different geographical ranges and lastly according very numerous experiments made during several years that most careful observer gärtner they can crossed only with much difficulty could hardly wish for better evidence the two forms being specifically distinct the other hand they are united many intermediate links and very doubtful whether these links are hybrids and there seems overwhelming amount experimental evidence showing that they descend from common parents and consequently must ranked varieties close investigation most cases will bring naturalists agreement how rank doubtful forms yet must confessed that the best known countries that find the greatest number forms doubtful value have been struck with the fact that any animal plant state nature highly useful man from any cause closely attract his attention varieties will almost universally found recorded these varieties moreover will often ranked some authors species look the common oak how closely has been studied yet german author makes more than dozen species out forms which are very generally considered varieties and this country the highest botanical authorities and practical men can quoted show that the sessile and pedunculated oaks are either good and distinct species mere varieties when young naturalist commences the study group organisms quite unknown him first much perplexed determine what differences consider specific and what varieties for knows nothing the amount and kind variation which the group subject and this shows least how very generally there some variation but confine his attention one class within one country will soon make his mind how rank most the doubtful forms his general tendency will make many species for will become impressed just like the pigeon poultry fancier before alluded with the amount difference the forms which continually studying and has little general knowledge analogical variation other groups and other countries which correct his first impressions extends the range his observations will meet with more cases difficulty for will encounter greater number closely allied forms but his observations widely extended will the end generally enabled make his own mind which call varieties and which species but will succeed this the expense admitting much variation and the truth this admission will often disputed other naturalists when moreover comes study allied forms brought from countries not now continuous which case can hardly hope find the intermediate links between his doubtful forms will have trust almost entirely analogy and his difficulties will rise climax certainly clear line demarcation has yet been drawn between species and sub species that the forms which the opinion some naturalists come very near but not quite arrive the rank species again between sub species and well marked varieties between lesser varieties and individual differences these differences blend into each other insensible series and series impresses the mind with the idea actual passage hence look individual differences though small interest the systematist high importance for being the first step towards such slight varieties are barely thought worth recording works natural history and look varieties which are any degree more distinct and permanent steps leading more strongly marked and more permanent varieties and these latter leading sub species and species the passage from one stage difference another and higher stage may some cases due merely the long continued action different physical conditions two different regions but have not much faith this view and attribute the passage variety from state which differs very slightly from its parent one which differs more the action natural selection accumulating will hereafter more fully explained differences structure certain definite directions hence believe well marked variety may justly called incipient species but whether this belief justifiable must judged the general weight the several facts and views given throughout this work need not supposed that all varieties incipient species necessarily attain the rank species they may whilst this incipient state become extinct they may endure varieties for very long periods has been shown the case wollaston with the varieties certain fossil land shells madeira variety were flourish exceed numbers the parent species would then rank the species and the species the variety might come supplant and exterminate the parent species both might exist and both rank independent species but shall hereafter have return this subject from these remarks will seen that look the term species one arbitrarily given for the sake convenience set individuals closely resembling each other and that does not essentially differ from the term variety which given less distinct and more fluctuating forms the term variety again comparison with mere individual differences also applied arbitrarily and for mere convenience sake guided theoretical considerations thought that some interesting results might obtained regard the nature and relations the species which vary most tabulating all the varieties several well worked floras first this seemed simple task but watson whom much indebted for valuable advice and assistance this subject soon convinced that there were many difficulties did subsequently hooker even stronger terms shall reserve for future work the discussion these difficulties and the tables themselves the proportional numbers the varying species hooker permits add that after having carefully read manuscript and examined the tables thinks that the following statements are fairly well established the whole subject however treated necessarily here with much brevity rather perplexing and allusions cannot avoided the struggle for existence divergence character and other questions hereafter discussed alph candolle and others have shown that plants which have very wide ranges generally present varieties and this might have been expected they become exposed diverse physical conditions and they come into competition which shall hereafter see far more important circumstance with different sets organic beings but tables further show that any limited country the species which are most common that abound most individuals and the species which are most widely diffused within their own country and this different consideration from wide range and certain extent from commonness often give rise varieties sufficiently well marked have been recorded botanical works hence the most flourishing they may called the dominant species those which range widely over the world are the most diffused their own country and are the most numerous individuals which oftenest produce well marked varieties consider them incipient species and this perhaps might have been anticipated for varieties order become any degree permanent necessarily have struggle with the other inhabitants the country the species which are already dominant will the most likely yield offspring which though some slight degree modified will still inherit those advantages that enabled their parents become dominant over their compatriots the plants inhabiting country and described any flora divided into two equal masses all those the larger genera being placed one side and all those the smaller genera the other side somewhat larger number the very common and much diffused dominant species will found the side the larger genera this again might have been anticipated for the mere fact many species the same genus inhabiting any country shows that there something the organic inorganic conditions that country favourable the genus and consequently might have expected have found the larger genera those including many species large proportional number dominant species but many causes tend obscure this result that surprised that tables show even small majority the side the larger genera will here allude only two causes obscurity fresh water and salt loving plants have generally very wide ranges and are much diffused but this seems connected with the nature the stations inhabited them and has little relation the size the genera which the species belong again plants low the scale organisation are generally much more widely diffused than plants higher the scale and here again there close relation the size the genera the cause lowly organised plants ranging widely will discussed our chapter geographical distribution from looking species only strongly marked and well defined varieties was led anticipate that the species the larger genera each country would oftener present varieties than the species the smaller genera for wherever many closely related species species the same genus have been formed many varieties incipient species ought general rule now forming where many large trees grow expect find saplings where many species genus have been formed through variation circumstances have been favourable for variation and hence might expect that the circumstances would generally still favourable variation the other hand look each species special act creation there apparent reason why more varieties should occur group having many species than one having few test the truth this anticipation have arranged the plants twelve countries and the coleopterous insects two districts into two nearly equal masses the species the larger genera one side and those the smaller genera the other side and has invariably proved the case that larger proportion the species the side the larger genera present varieties than the side the smaller genera moreover the species the large genera which present any varieties invariably present larger average number varieties than the species the small genera both these results follow when another division made and when all the smallest genera with from only one four species are absolutely excluded from the tables these facts are plain signification the view that species are only strongly marked and permanent varieties for wherever many species the same genus have been formed where may use the expression the manufactory species has been active ought generally find the manufactory still action more especially have every reason believe the process manufacturing new species slow one and this certainly the case varieties looked incipient species for tables clearly show general rule that wherever many species genus have been formed the species that genus present number varieties that incipient species beyond the average not that all large genera are now varying much and are thus increasing the number their species that small genera are now varying and increasing for this had been would have been fatal theory inasmuch geology plainly tells that small genera have the lapse time often increased greatly size and that large genera have often come their maxima declined and disappeared all that want show that where many species genus have been formed average many are still forming and this holds good there are other relations between the species large genera and their recorded varieties which deserve notice have seen that there infallible criterion which distinguish species and well marked varieties and those cases which intermediate links have not been found between doubtful forms naturalists are compelled come determination the amount difference between them judging analogy whether not the amount suffices raise one both the rank species hence the amount difference one very important criterion settling whether two forms should ranked species varieties now fries has remarked regard plants and westwood regard insects that large genera the amount difference between the species often exceedingly small have endeavoured test this numerically averages and far imperfect results they always confirm the view have also consulted some sagacious and most experienced observers and after deliberation they concur this view this respect therefore the species the larger genera resemble varieties more than the species the smaller genera the case may put another way and may said that the larger genera which number varieties incipient species greater than the average are now manufacturing many the species already manufactured still certain extent resemble varieties for they differ from each other less than usual amount difference moreover the species the large genera are related each other the same manner the varieties any one species are related each other naturalist pretends that all the species genus are equally distinct from each other they may generally divided into sub genera sections lesser groups fries has well remarked little groups species are generally clustered like satellites around certain other species and what are varieties but groups forms unequally related each other and clustered round certain forms that round their parent species undoubtedly there one most important point difference between varieties and species namely that the amount difference between varieties when compared with each other with their parent species much less than that between the species the same genus but when come discuss the principle call divergence character shall see how this may explained and how the lesser differences between varieties will tend increase into the greater differences between species there one other point which seems worth notice varieties generally have much restricted ranges this statement indeed scarcely more than truism for variety were found have wider range than that its supposed parent species their denominations ought reversed but there also reason believe that those species which are very closely allied other species and far resemble varieties often have much restricted ranges for instance watson has marked for the well sifted london catalogue plants 4th edition plants which are therein ranked species but which considers closely allied other species doubtful value these reputed species range average over the provinces into which watson has divided great britain now this same catalogue acknowledged varieties are recorded and these range over provinces whereas the species which these varieties belong range over provinces that the acknowledged varieties have very nearly the same restricted average range have those very closely allied forms marked for watson doubtful species but which are almost universally ranked british botanists good and true species finally then varieties have the same general characters species for they cannot distinguished from species except firstly the discovery intermediate linking forms and the occurrence such links cannot affect the actual characters the forms which they connect and except secondly certain amount difference for two forms differing very little are generally ranked varieties notwithstanding that intermediate linking forms have not been discovered but the amount difference considered necessary give two forms the rank species quite indefinite genera having more than the average number species any country the species these genera have more than the average number varieties large genera the species are apt closely but unequally allied together forming little clusters round certain species species very closely allied other species apparently have restricted ranges all these several respects the species large genera present strong analogy with varieties and can clearly understand these analogies species have once existed varieties and have thus originated whereas these analogies are utterly inexplicable each species has been independently created have also seen that the most flourishing and dominant species the larger genera which average vary most and varieties shall hereafter see tend become converted into new and distinct species the larger genera thus tend become larger and throughout nature the forms life which are now dominant tend become still more dominant leaving many modified and dominant descendants but steps hereafter explained the larger genera also tend break into smaller genera and thus the forms life throughout the universe become divided into groups subordinate groups chapter iii struggle for existence bears natural selection the term used wide sense geometrical powers increase rapid increase naturalised animals and plants nature the checks increase competition universal effects climate protection from the number individuals complex relations all animals and plants throughout nature struggle for life most severe between individuals and varieties the same species often severe between species the same genus the relation organism organism the most important all relations before entering the subject this chapter must make few preliminary remarks show how the struggle for existence bears natural selection has been seen the last chapter that amongst organic beings state nature there some individual variability indeed not aware that this has ever been disputed immaterial for whether multitude doubtful forms called species sub species varieties what rank for instance the two three hundred doubtful forms british plants are entitled hold the existence any well marked varieties admitted but the mere existence individual variability and some few well marked varieties though necessary the foundation for the work helps but little understanding how species arise nature how have all those exquisite adaptations one part the organisation another part and the conditions life and one distinct organic being another being been perfected see these beautiful adaptations most plainly the woodpecker and missletoe and only little less plainly the humblest parasite which clings the hairs quadruped feathers bird the structure the beetle which dives through the water the plumed seed which wafted the gentlest breeze short see beautiful adaptations everywhere and every part the organic world again may asked how that varieties which have called incipient species become ultimately converted into good and distinct species which most cases obviously differ from each other far more than the varieties the same species how those groups species which constitute what are called distinct genera and which differ from each other more than the species the same genus arise all these results shall more fully see the next chapter follow inevitably from the struggle for life owing this struggle for life any variation however slight and from whatever cause proceeding any degree profitable individual any species its infinitely complex relations other organic beings and external nature will tend the preservation that individual and will generally inherited its offspring the offspring also will thus have better chance surviving for the many individuals any species which are periodically born but small number can survive have called this principle which each slight variation useful preserved the term natural selection order mark its relation man power selection have seen that man selection can certainly produce great results and can adapt organic beings his own uses through the accumulation slight but useful variations given him the hand nature but natural selection shall hereafter see power incessantly ready for action and immeasurably superior man feeble efforts the works nature are those art will now discuss little more detail the struggle for existence future work this subject shall treated well deserves much greater length the elder candolle and lyell have largely and philosophically shown that all organic beings are exposed severe competition regard plants one has treated this subject with more spirit and ability than herbert dean manchester evidently the result his great horticultural knowledge nothing easier than admit words the truth the universal struggle for life more difficult least have found than constantly bear this conclusion mind yet unless thoroughly engrained the mind convinced that the whole economy nature with every fact distribution rarity abundance extinction and variation will dimly seen quite misunderstood behold the face nature bright with gladness often see superabundance food not see forget that the birds which are idly singing round mostly live insects seeds and are thus constantly destroying life forget how largely these songsters their eggs their nestlings are destroyed birds and beasts prey not always bear mind that though food may now superabundant not all seasons each recurring year should premise that use the term struggle for existence large and metaphorical sense including dependence one being another and including which more important not only the life the individual but success leaving progeny two canine animals time dearth may truly said struggle with each other which shall get food and live but plant the edge desert said struggle for life against the drought though more properly should said dependent the moisture plant which annually produces thousand seeds which average only one comes maturity may more truly said struggle with the plants the same and other kinds which already clothe the ground the missletoe dependent the apple and few other trees but can only far fetched sense said struggle with these trees for too many these parasites grow the same tree will languish and die but several seedling missletoes growing close together the same branch may more truly said struggle with each other the missletoe disseminated birds its existence depends birds and may metaphorically said struggle with other fruit bearing plants order tempt birds devour and thus disseminate its seeds rather than those other plants these several senses which pass into each other use for convenience sake the general term struggle for existence struggle for existence inevitably follows from the high rate which all organic beings tend increase every being which during its natural lifetime produces several eggs seeds must suffer destruction during some period its life and during some season occasional year otherwise the principle geometrical increase its numbers would quickly become inordinately great that country could support the product hence more individuals are produced than can possibly survive there must every case struggle for existence either one individual with another the same species with the individuals distinct species with the physical conditions life the doctrine malthus applied with manifold force the whole animal and vegetable kingdoms for this case there can artificial increase food and prudential restraint from marriage although some species may now increasing more less rapidly numbers all cannot for the world would not hold them there exception the rule that every organic being naturally increases high rate that not destroyed the earth would soon covered the progeny single pair even slow breeding man has doubled twenty five years and this rate few thousand years there would literally not standing room for his progeny linnæus has calculated that annual plant produced only two seeds and there plant unproductive this and their seedlings next year produced two and then twenty years there would million plants the elephant reckoned the slowest breeder all known animals and have taken some pains estimate its probable minimum rate natural increase will under the mark assume that breeds when thirty years old and goes breeding till ninety years old bringing forth three pair young this interval this the end the fifth century there would alive fifteen million elephants descended from the first pair but have better evidence this subject than mere theoretical calculations namely the numerous recorded cases the astonishingly rapid increase various animals state nature when circumstances have been favourable them during two three following seasons still more striking the evidence from our domestic animals many kinds which have run wild several parts the world the statements the rate increase slow breeding cattle and horses south america and latterly australia had not been well authenticated they would have been quite incredible with plants cases could given introduced plants which have become common throughout whole islands period less than ten years several the plants now most numerous over the wide plains plata clothing square leagues surface almost the exclusion all other plants have been introduced from europe and there are plants which now range india hear from falconer from cape comorin the himalaya which have been imported from america since its discovery such cases and endless instances could given one supposes that the fertility these animals plants has been suddenly and temporarily increased any sensible degree the obvious explanation that the conditions life have been very favourable and that there has consequently been less destruction the old and young and that nearly all the young have been enabled breed such cases the geometrical ratio increase the result which never fails surprising simply explains the extraordinarily rapid increase and wide diffusion naturalised productions their new homes state nature almost every plant produces seed and amongst animals there are very few which not annually pair hence may confidently assert that all plants and animals are tending increase geometrical ratio that all would most rapidly stock every station which they could any how exist and that the geometrical tendency increase must checked destruction some period life our familiarity with the larger domestic animals tends think mislead see great destruction falling them and forget that thousands are annually slaughtered for food and that state nature equal number would have somehow disposed the only difference between organisms which annually produce eggs seeds the thousand and those which produce extremely few that the slow breeders would require few more years people under favourable conditions whole district let ever large the condor lays couple eggs and the ostrich score and yet the same country the condor may the more numerous the two the fulmar petrel lays but one egg yet believed the most numerous bird the world one fly deposits hundreds eggs and another like the hippobosca single one but this difference does not determine how many individuals the two species can supported district large number eggs some importance those species which depend rapidly fluctuating amount food for allows them rapidly increase number but the real importance large number eggs seeds make for much destruction some period life and this period the great majority cases early one animal can any way protect its own eggs young small number may produced and yet the average stock fully kept but many eggs young are destroyed many must produced the species will become extinct would suffice keep the full number tree which lived average for thousand years single seed were produced once thousand years supposing that this seed were never destroyed and could ensured germinate fitting place that all cases the average number any animal plant depends only indirectly the number its eggs seeds looking nature most necessary keep the foregoing considerations always mind never forget that every single organic being around may said striving the utmost increase numbers that each lives struggle some period its life that heavy destruction inevitably falls either the young old during each generation recurrent intervals lighten any check mitigate the destruction ever little and the number the species will almost instantaneously increase any amount the face nature may compared yielding surface with ten thousand sharp wedges packed close together and driven inwards incessant blows sometimes one wedge being struck and then another with greater force what checks the natural tendency each species increase number most obscure look the most vigorous species much swarms numbers much will its tendency increase still further increased know not exactly what the checks are even one single instance nor will this surprise any one who reflects how ignorant are this head even regard mankind incomparably better known than any other animal this subject has been ably treated several authors and shall future work discuss some the checks considerable length more especially regard the feral animals south america here will make only few remarks just recall the reader mind some the chief points eggs very young animals seem generally suffer most but this not invariably the case with plants there vast destruction seeds but from some observations which have made believe that the seedlings which suffer most from germinating ground already thickly stocked with other plants seedlings also are destroyed vast numbers various enemies for instance piece ground three feet long and two wide dug and cleared and where there could choking from other plants marked all the seedlings our native weeds they came and out the 357 less than 295 were destroyed chiefly slugs and insects turf which has long been mown and the case would the same with turf closely browsed quadrupeds let grow the more vigorous plants gradually kill the less vigorous though fully grown plants thus out twenty species growing little plot turf three feet four nine species perished from the other species being allowed grow freely the amount food for each species course gives the extreme limit which each can increase but very frequently not the obtaining food but the serving prey other animals which determines the average numbers species thus there seems little doubt that the stock partridges grouse and hares any large estate depends chiefly the destruction vermin not one head game were shot during the next twenty years england and the same time vermin were destroyed there would all probability less game than present although hundreds thousands game animals are now annually killed the other hand some cases with the elephant and rhinoceros none are destroyed beasts prey even the tiger india most rarely dares attack young elephant protected its dam climate plays important part determining the average numbers species and periodical seasons extreme cold drought believe the most effective all checks estimated that the winter 1854 destroyed four fifths the birds own grounds and this tremendous destruction when remember that ten per cent extraordinarily severe mortality from epidemics with man the action climate seems first sight quite independent the struggle for existence but far climate chiefly acts reducing food brings the most severe struggle between the individuals whether the same distinct species which subsist the same kind food even when climate for instance extreme cold acts directly will the least vigorous those which have got least food through the advancing winter which will suffer most when travel from south north from damp region dry invariably see some species gradually getting rarer and rarer and finally disappearing and the change climate being conspicuous are tempted attribute the whole effect its direct action but this very false view forget that each species even where most abounds constantly suffering enormous destruction some period its life from enemies from competitors for the same place and food and these enemies competitors the least degree favoured any slight change climate they will increase numbers and each area already fully stocked with inhabitants the other species will decrease when travel southward and see species decreasing numbers may feel sure that the cause lies quite much other species being favoured this one being hurt when travel northward but somewhat lesser degree for the number species all kinds and therefore competitors decreases northwards hence going northward ascending mountain far oftener meet with stunted forms due the directly injurious action climate than proceeding southwards descending mountain when reach the arctic regions snow capped summits absolute deserts the struggle for life almost exclusively with the elements that climate acts main part indirectly favouring other species may clearly see the prodigious number plants our gardens which can perfectly well endure our climate but which never become naturalised for they cannot compete with our native plants nor resist destruction our native animals when species owing highly favourable circumstances increases inordinately numbers small tract epidemics least this seems generally occur with our game animals often ensue and here have limiting check independent the struggle for life but even some these called epidemics appear due parasitic worms which have from some cause possibly part through facility diffusion amongst the crowded animals been disproportionably favoured and here comes sort struggle between the parasite and its prey the other hand many cases large stock individuals the same species relatively the numbers its enemies absolutely necessary for its preservation thus can easily raise plenty corn and rape seed etc our fields because the seeds are great excess compared with the number birds which feed them nor can the birds though having superabundance food this one season increase number proportionally the supply seed their numbers are checked during winter but any one who has tried knows how troublesome get seed from few wheat other such plants garden have this case lost every single seed this view the necessity large stock the same species for its preservation explains believe some singular facts nature such that very rare plants being sometimes extremely abundant the few spots where they occur and that some social plants being social that abounding individuals even the extreme confines their range for such cases may believe that plant could exist only where the conditions its life were favourable that many could exist together and thus save each other from utter destruction should add that the good effects frequent intercrossing and the ill effects close interbreeding probably come into play some these cases but this intricate subject will not here enlarge many cases are record showing how complex and unexpected are the checks and relations between organic beings which have struggle together the same country will give only single instance which though simple one has interested staffordshire the estate relation where had ample means investigation there was large and extremely barren heath which had never been touched the hand man but several hundred acres exactly the same nature had been enclosed twenty five years previously and planted with scotch fir the change the native vegetation the planted part the heath was most remarkable more than generally seen passing from one quite different soil another not only the proportional numbers the heath plants were wholly changed but twelve species plants not counting grasses and carices flourished the plantations which could not found the heath the effect the insects must have been still greater for six insectivorous birds were very common the plantations which were not seen the heath and the heath was frequented two three distinct insectivorous birds here see how potent has been the effect the introduction single tree nothing whatever else having been done with the exception that the land had been enclosed that cattle could not enter but how important element enclosure plainly saw near farnham surrey here there are extensive heaths with few clumps old scotch firs the distant hill tops within the last ten years large spaces have been enclosed and self sown firs are now springing multitudes close together that all cannot live when ascertained that these young trees had not been sown planted was much surprised their numbers that went several points view whence could examine hundreds acres the unenclosed heath and literally could not see single scotch fir except the old planted clumps but looking closely between the stems the heath found multitude seedlings and little trees which had been perpetually browsed down the cattle one square yard point some hundred yards distant from one the old clumps counted thirty two little trees and one them judging from the rings growth had during twenty six years tried raise its head above the stems the heath and had failed wonder that soon the land was enclosed became thickly clothed with vigorously growing young firs yet the heath was extremely barren and extensive that one would ever have imagined that cattle would have closely and effectually searched for food here see that cattle absolutely determine the existence the scotch fir but several parts the world insects determine the existence cattle perhaps paraguay offers the most curious instance this for here neither cattle nor horses nor dogs have ever run wild though they swarm southward and northward feral state and azara and rengger have shown that this caused the greater number paraguay certain fly which lays its eggs the navels these animals when first born the increase these flies numerous they are must habitually checked some means probably birds hence certain insectivorous birds whose numbers are probably regulated hawks beasts prey were increase paraguay the flies would decrease then cattle and horses would become feral and this would certainly greatly alter indeed have observed parts south america the vegetation this again would largely affect the insects and this just have seen staffordshire the insectivorous birds and onwards ever increasing circles complexity began this series insectivorous birds and have ended with them not that nature the relations can ever simple this battle within battle must ever recurring with varying success and yet the long run the forces are nicely balanced that the face nature remains uniform for long periods time though assuredly the merest trifle would often give the victory one organic being over another nevertheless profound our ignorance and high our presumption that marvel when hear the extinction organic being and not see the cause invoke cataclysms desolate the world invent laws the duration the forms life tempted give one more instance showing how plants and animals most remote the scale nature are bound together web complex relations shall hereafter have occasion show that the exotic lobelia fulgens this part england never visited insects and consequently from its peculiar structure never can set seed many our orchidaceous plants absolutely require the visits moths remove their pollen masses and thus fertilise them have also reason believe that humble bees are indispensable the fertilisation the heartsease viola tricolor for other bees not visit this flower from experiments which have tried have found that the visits bees not indispensable are least highly beneficial the fertilisation our clovers but humble bees alone visit the common red clover trifolium pratense other bees cannot reach the nectar hence have very little doubt that the whole genus humble bees became extinct very rare england the heartsease and red clover would become very rare wholly disappear the number humble bees any district depends great degree the number field mice which destroy their combs and nests and newman who has long attended the habits humble bees believes that more than two thirds them are thus destroyed all over england now the number mice largely dependent every one knows the number cats and newman says near villages and small towns have found the nests humble bees more numerous than elsewhere which attribute the number cats that destroy the mice hence quite credible that the presence feline animal large numbers district might determine through the intervention first mice and then bees the frequency certain flowers that district the case every species many different checks acting different periods life and during different seasons years probably come into play some one check some few being generally the most potent but all concurring determining the average number even the existence the species some cases can shown that widely different checks act the same species different districts when look the plants and bushes clothing entangled bank are tempted attribute their proportional numbers and kinds what call chance but how false view this every one has heard that when american forest cut down very different vegetation springs but has been observed that the trees now growing the ancient indian mounds the southern united states display the same beautiful diversity and proportion kinds the surrounding virgin forests what struggle between the several kinds trees must here have gone during long centuries each annually scattering its seeds the thousand what war between insect and insect between insects snails and other animals with birds and beasts prey all striving increase and all feeding each other the trees their seeds and seedlings the other plants which first clothed the ground and thus checked the growth the trees throw handful feathers and all must fall the ground according definite laws but how simple this problem compared the action and reaction the innumerable plants and animals which have determined the course centuries the proportional numbers and kinds trees now growing the old indian ruins the dependency one organic being another parasite its prey lies generally between beings remote the scale nature this often the case with those which may strictly said struggle with each other for existence the case locusts and grass feeding quadrupeds but the struggle almost invariably will most severe between the individuals the same species for they frequent the same districts require the same food and are exposed the same dangers the case varieties the same species the struggle will generally almost equally severe and sometimes see the contest soon decided for instance several varieties wheat sown together and the mixed seed resown some the varieties which best suit the soil climate are naturally the most fertile will beat the others and yield more seed and will consequently few years quite supplant the other varieties keep mixed stock even such extremely close varieties the variously coloured sweet peas they must each year harvested separately and the seed then mixed due proportion otherwise the weaker kinds will steadily decrease numbers and disappear again with the varieties sheep has been asserted that certain mountain varieties will starve out other mountain varieties that they cannot kept together the same result has followed from keeping together different varieties the medicinal leech may even doubted whether the varieties any one our domestic plants animals have exactly the same strength habits and constitution that the original proportions mixed stock could kept for half dozen generations they were allowed struggle together like beings state nature and the seed young were not annually sorted species the same genus have usually though means invariably some similarity habits and constitution and always structure the struggle will generally more severe between species the same genus when they come into competition with each other than between species distinct genera see this the recent extension over parts the united states one species swallow having caused the decrease another species the recent increase the missel thrush parts scotland has caused the decrease the song thrush how frequently hear one species rat taking the place another species under the most different climates russia the small asiatic cockroach has everywhere driven before its great congener one species charlock will supplant another and other cases can dimly see why the competition should most severe between allied forms which fill nearly the same place the economy nature but probably one case could precisely say why one species has been victorious over another the great battle life corollary the highest importance may deduced from the foregoing remarks namely that the structure every organic being related the most essential yet often hidden manner that all other organic beings with which comes into competition for food residence from which has escape which preys this obvious the structure the teeth and talons the tiger and that the legs and claws the parasite which clings the hair the tiger body but the beautifully plumed seed the dandelion and the flattened and fringed legs the water beetle the relation seems first confined the elements air and water yet the advantage plumed seeds doubt stands the closest relation the land being already thickly clothed other plants that the seeds may widely distributed and fall unoccupied ground the water beetle the structure its legs well adapted for diving allows compete with other aquatic insects hunt for its own prey and escape serving prey other animals the store nutriment laid within the seeds many plants seems first sight have sort relation other plants but from the strong growth young plants produced from such seeds peas and beans when sown the midst long grass suspect that the chief use the nutriment the seed favour the growth the young seedling whilst struggling with other plants growing vigorously all around look plant the midst its range why does not double quadruple its numbers know that can perfectly well withstand little more heat cold dampness dryness for elsewhere ranges into slightly hotter colder damper drier districts this case can clearly see that wished imagination give the plant the power increasing number should have give some advantage over its competitors over the animals which preyed the confines its geographical range change constitution with respect climate would clearly advantage our plant but have reason believe that only few plants animals range far that they are destroyed the rigour the climate alone not until reach the extreme confines life the arctic regions the borders utter desert will competition cease the land may extremely cold dry yet there will competition between some few species between the individuals the same species for the warmest dampest spots hence also can see that when plant animal placed new country amongst new competitors though the climate may exactly the same its former home yet the conditions its life will generally changed essential manner wished increase its average numbers its new home should have modify different way what should have done its native country for should have give some advantage over different set competitors enemies good thus try our imagination give any form some advantage over another probably single instance should know what succeed will convince our ignorance the mutual relations all organic beings conviction necessary seems difficult acquire all that can keep steadily mind that each organic being striving increase geometrical ratio that each some period its life during some season the year during each generation intervals has struggle for life and suffer great destruction when reflect this struggle may console ourselves with the full belief that the war nature not incessant that fear felt that death generally prompt and that the vigorous the healthy and the happy survive and multiply chapter natural selection natural selection its power compared with man selection its power characters trifling importance its power all ages and both sexes sexual selection the generality intercrosses between individuals the same species circumstances favourable and unfavourable natural selection namely intercrossing isolation number individuals slow action extinction caused natural selection divergence character related the diversity inhabitants any small area and naturalisation action natural selection through divergence character and extinction the descendants from common parent explains the grouping all organic beings how will the struggle for existence discussed too briefly the last chapter act regard variation can the principle selection which have seen potent the hands man apply nature think shall see that can act most effectually let borne mind what endless number strange peculiarities our domestic productions and lesser degree those under nature vary and how strong the hereditary tendency under domestication may truly said that the whole organisation becomes some degree plastic let borne mind how infinitely complex and close fitting are the mutual relations all organic beings each other and their physical conditions life can then thought improbable seeing that variations useful man have undoubtedly occurred that other variations useful some way each being the great and complex battle life should sometimes occur the course thousands generations such occur can doubt remembering that many more individuals are born than can possibly survive that individuals having any advantage however slight over others would have the best chance surviving and procreating their kind the other hand may feel sure that any variation the least degree injurious would rigidly destroyed this preservation favourable variations and the rejection injurious variations call natural selection variations neither useful nor injurious would not affected natural selection and would left fluctuating element perhaps see the species called polymorphic shall best understand the probable course natural selection taking the case country undergoing some physical change for instance climate the proportional numbers its inhabitants would almost immediately undergo change and some species might become extinct may conclude from what have seen the intimate and complex manner which the inhabitants each country are bound together that any change the numerical proportions some the inhabitants independently the change climate itself would most seriously affect many the others the country were open its borders new forms would certainly immigrate and this also would seriously disturb the relations some the former inhabitants let remembered how powerful the influence single introduced tree mammal has been shown but the case island country partly surrounded barriers into which new and better adapted forms could not freely enter should then have places the economy nature which would assuredly better filled some the original inhabitants were some manner modified for had the area been open immigration these same places would have been seized intruders such case every slight modification which the course ages chanced arise and which any way favoured the individuals any the species better adapting them their altered conditions would tend preserved and natural selection would thus have free scope for the work improvement have reason believe stated the first chapter that change the conditions life specially acting the reproductive system causes increases variability and the foregoing case the conditions life are supposed have undergone change and this would manifestly favourable natural selection giving better chance profitable variations occurring and unless profitable variations occur natural selection can nothing not that believe any extreme amount variability necessary man can certainly produce great results adding any given direction mere individual differences could nature but far more easily from having incomparably longer time her disposal nor believe that any great physical change climate any unusual degree isolation check immigration actually necessary produce new and unoccupied places for natural selection fill modifying and improving some the varying inhabitants for all the inhabitants each country are struggling together with nicely balanced forces extremely slight modifications the structure habits one inhabitant would often give advantage over others and still further modifications the same kind would often still further increase the advantage country can named which all the native inhabitants are now perfectly adapted each other and the physical conditions under which they live that none them could anyhow improved for all countries the natives have been far conquered naturalised productions that they have allowed foreigners take firm possession the land and foreigners have thus everywhere beaten some the natives may safely conclude that the natives might have been modified with advantage have better resisted such intruders man can produce and certainly has produced great result his methodical and unconscious means selection what may not nature effect man can act only external and visible characters nature cares nothing for appearances except far they may useful any being she can act every internal organ every shade constitutional difference the whole machinery life man selects only for his own good nature only for that the being which she tends every selected character fully exercised her and the being placed under well suited conditions life man keeps the natives many climates the same country seldom exercises each selected character some peculiar and fitting manner feeds long and short beaked pigeon the same food does not exercise long backed long legged quadruped any peculiar manner exposes sheep with long and short wool the same climate does not allow the most vigorous males struggle for the females does not rigidly destroy all inferior animals but protects during each varying season far lies his power all his productions often begins his selection some half monstrous form least some modification prominent enough catch his eye plainly useful him under nature the slightest difference structure constitution may well turn the nicely balanced scale the struggle for life and preserved how fleeting are the wishes and efforts man how short his time and consequently how poor will his products compared with those accumulated nature during whole geological periods can wonder then that nature productions should far truer character than man productions that they should infinitely better adapted the most complex conditions life and should plainly bear the stamp far higher workmanship may said that natural selection daily and hourly scrutinising throughout the world every variation even the slightest rejecting that which bad preserving and adding all that good silently and insensibly working whenever and wherever opportunity offers the improvement each organic being relation its organic and inorganic conditions life see nothing these slow changes progress until the hand time has marked the long lapse ages and then imperfect our view into long past geological ages that only see that the forms life are now different from what they formerly were although natural selection can act only through and for the good each being yet characters and structures which are apt consider very trifling importance may thus acted when see leaf eating insects green and bark feeders mottled grey the alpine ptarmigan white winter the red grouse the colour heather and the black grouse that peaty earth must believe that these tints are service these birds and insects preserving them from danger grouse not destroyed some period their lives would increase countless numbers they are known suffer largely from birds prey and hawks are guided eyesight their prey much that parts the continent persons are warned not keep white pigeons being the most liable destruction hence can see reason doubt that natural selection might most effective giving the proper colour each kind grouse and keeping that colour when once acquired true and constant nor ought think that the occasional destruction animal any particular colour would produce little effect should remember how essential flock white sheep destroy every lamb with the faintest trace black plants the down the fruit and the colour the flesh are considered botanists characters the most trifling importance yet hear from excellent horticulturist downing that the united states smooth skinned fruits suffer far more from beetle curculio than those with down that purple plums suffer far more from certain disease than yellow plums whereas another disease attacks yellow fleshed peaches far more than those with other coloured flesh with all the aids art these slight differences make great difference cultivating the several varieties assuredly state nature where the trees would have struggle with other trees and with host enemies such differences would effectually settle which variety whether smooth downy yellow purple fleshed fruit should succeed looking many small points difference between species which far our ignorance permits judge seem quite unimportant must not forget that climate food etc probably produce some slight and direct effect however far more necessary bear mind that there are many unknown laws correlation growth which when one part the organisation modified through variation and the modifications are accumulated natural selection for the good the being will cause other modifications often the most unexpected nature see that those variations which under domestication appear any particular period life tend reappear the offspring the same period for instance the seeds the many varieties our culinary and agricultural plants the caterpillar and cocoon stages the varieties the silkworm the eggs poultry and the colour the down their chickens the horns our sheep and cattle when nearly adult state nature natural selection will enabled act and modify organic beings any age the accumulation profitable variations that age and their inheritance corresponding age profit plant have its seeds more and more widely disseminated the wind can see greater difficulty this being effected through natural selection than the cotton planter increasing and improving selection the down the pods his cotton trees natural selection may modify and adapt the larva insect score contingencies wholly different from those which concern the mature insect these modifications will doubt affect through the laws correlation the structure the adult and probably the case those insects which live only for few hours and which never feed large part their structure merely the correlated result successive changes the structure their larvæ conversely modifications the adult will probably often affect the structure the larva but all cases natural selection will ensure that modifications consequent other modifications different period life shall not the least degree injurious for they became they would cause the extinction the species natural selection will modify the structure the young relation the parent and the parent relation the young social animals will adapt the structure each individual for the benefit the community each consequence profits the selected change what natural selection cannot modify the structure one species without giving any advantage for the good another species and though statements this effect may found works natural history cannot find one case which will bear investigation structure used only once animal whole life high importance might modified any extent natural selection for instance the great jaws possessed certain insects and used exclusively for opening the cocoon the hard tip the beak nestling birds used for breaking the egg has been asserted that the best short beaked tumbler pigeons more perish the egg than are able get out that fanciers assist the act hatching now nature had make the beak full grown pigeon very short for the bird own advantage the process modification would very slow and there would simultaneously the most rigorous selection the young birds within the egg which had the most powerful and hardest beaks for all with weak beaks would inevitably perish more delicate and more easily broken shells might selected the thickness the shell being known vary like every other structure sexual selection inasmuch peculiarities often appear under domestication one sex and become hereditarily attached that sex the same fact probably occurs under nature and natural selection will able modify one sex its functional relations the other sex relation wholly different habits life the two sexes sometimes the case with insects and this leads say few words what call sexual selection this depends not struggle for existence but struggle between the males for possession the females the result not death the unsuccessful competitor but few offspring sexual selection therefore less rigorous than natural selection generally the most vigorous males those which are best fitted for their places nature will leave most progeny but many cases victory will depend not general vigour but having special weapons confined the male sex hornless stag spurless cock would have poor chance leaving offspring sexual selection always allowing the victor breed might surely give indomitable courage length the spur and strength the wing strike the spurred leg well the brutal cock fighter who knows well that can improve his breed careful selection the best cocks how low the scale nature this law battle descends know not male alligators have been described fighting bellowing and whirling round like indians war dance for the possession the females male salmons have been seen fighting all day long male stag beetles often bear wounds from the huge mandibles other males the war perhaps severest between the males polygamous animals and these seem oftenest provided with special weapons the males carnivorous animals are already well armed though them and others special means defence may given through means sexual selection the mane the lion the shoulder pad the boar and the hooked jaw the male salmon for the shield may important for victory the sword spear amongst birds the contest often more peaceful character all those who have attended the subject believe that there the severest rivalry between the males many species attract singing the females the rock thrush guiana birds paradise and some others congregate and successive males display their gorgeous plumage and perform strange antics before the females which standing spectators last choose the most attractive partner those who have closely attended birds confinement well know that they often take individual preferences and dislikes thus sir heron has described how one pied peacock was eminently attractive all his hen birds may appear childish attribute any effect such apparently weak means cannot here enter the details necessary support this view but man can short time give elegant carriage and beauty his bantams according his standard beauty can see good reason doubt that female birds selecting during thousands generations the most melodious beautiful males according their standard beauty might produce marked effect strongly suspect that some well known laws with respect the plumage male and female birds comparison with the plumage the young can explained the view plumage having been chiefly modified sexual selection acting when the birds have come the breeding age during the breeding season the modifications thus produced being inherited corresponding ages seasons either the males alone the males and females but have not space here enter this subject thus believe that when the males and females any animal have the same general habits life but differ structure colour ornament such differences have been mainly caused sexual selection that individual males have had successive generations some slight advantage over other males their weapons means defence charms and have transmitted these advantages their male offspring yet would not wish attribute all such sexual differences this agency for see peculiarities arising and becoming attached the male sex our domestic animals the wattle male carriers horn like protuberances the cocks certain fowls etc which cannot believe either useful the males battle attractive the females see analogous cases under nature for instance the tuft hair the breast the turkey cock which can hardly either useful ornamental this bird indeed had the tuft appeared under domestication would have been called monstrosity illustrations the action natural selection order make clear how believe natural selection acts must beg permission give one two imaginary illustrations let take the case wolf which preys various animals securing some craft some strength and some fleetness and let suppose that the fleetest prey deer for instance had from any change the country increased numbers that other prey had decreased numbers during that season the year when the wolf hardest pressed for food can under such circumstances see reason doubt that the swiftest and slimmest wolves would have the best chance surviving and preserved selected provided always that they retained strength master their prey this some other period the year when they might compelled prey other animals can see more reason doubt this than that man can improve the fleetness his greyhounds careful and methodical selection that unconscious selection which results from each man trying keep the best dogs without any thought modifying the breed even without any change the proportional numbers the animals which our wolf preyed cub might born with innate tendency pursue certain kinds prey nor can this thought very improbable for often observe great differences the natural tendencies our domestic animals one cat for instance taking catch rats another mice one cat according john bringing home winged game another hares rabbits and another hunting marshy ground and almost nightly catching woodcocks snipes the tendency catch rats rather than mice known inherited now any slight innate change habit structure benefited individual wolf would have the best chance surviving and leaving offspring some its young would probably inherit the same habits structure and the repetition this process new variety might formed which would either supplant coexist with the parent form wolf again the wolves inhabiting mountainous district and those frequenting the lowlands would naturally forced hunt different prey and from the continued preservation the individuals best fitted for the two sites two varieties might slowly formed these varieties would cross and blend where they met but this subject intercrossing shall soon have return may add that according pierce there are two varieties the wolf inhabiting the catskill mountains the united states one with light greyhound like form which pursues deer and the other more bulky with shorter legs which more frequently attacks the shepherd flocks let now take more complex case certain plants excrete sweet juice apparently for the sake eliminating something injurious from their sap this effected glands the base the stipules some leguminosæ and the back the leaf the common laurel this juice though small quantity greedily sought insects let now suppose little sweet juice nectar excreted the inner bases the petals flower this case insects seeking the nectar would get dusted with pollen and would certainly often transport the pollen from one flower the stigma another flower the flowers two distinct individuals the same species would thus get crossed and the act crossing have good reason believe will hereafter more fully alluded would produce very vigorous seedlings which consequently would have the best chance flourishing and surviving some these seedlings would probably inherit the nectar excreting power those individual flowers which had the largest glands nectaries and which excreted most nectar would oftenest visited insects and would oftenest crossed and the long run would gain the upper hand those flowers also which had their stamens and pistils placed relation the size and habits the particular insects which visited them favour any degree the transportal their pollen from flower flower would likewise favoured selected might have taken the case insects visiting flowers for the sake collecting pollen instead nectar and pollen formed for the sole object fertilisation its destruction appears simple loss the plant yet little pollen were carried first occasionally and then habitually the pollen devouring insects from flower flower and cross thus effected although nine tenths the pollen were destroyed might still great gain the plant and those individuals which produced more and more pollen and had larger and larger anthers would selected when our plant this process the continued preservation natural selection more and more attractive flowers had been rendered highly attractive insects they would unintentionally their part regularly carry pollen from flower flower and that they can most effectually this could easily show many striking instances will give only one not very striking case but likewise illustrating one step the separation the sexes plants presently alluded some holly trees bear only male flowers which have four stamens producing rather small quantity pollen and rudimentary pistil other holly trees bear only female flowers these have full sized pistil and four stamens with shrivelled anthers which not grain pollen can detected having found female tree exactly sixty yards from male tree put the stigmas twenty flowers taken from different branches under the microscope and all without exception there were pollen grains and some profusion pollen the wind had set for several days from the female the male tree the pollen could not thus have been carried the weather had been cold and boisterous and therefore not favourable bees nevertheless every female flower which examined had been effectually fertilised the bees accidentally dusted with pollen having flown from tree tree search nectar but return our imaginary case soon the plant had been rendered highly attractive insects that pollen was regularly carried from flower flower another process might commence naturalist doubts the advantage what has been called the physiological division labour hence may believe that would advantageous plant produce stamens alone one flower one whole plant and pistils alone another flower another plant plants under culture and placed under new conditions life sometimes the male organs and sometimes the female organs become more less impotent now suppose this occur ever slight degree under nature then pollen already carried regularly from flower flower and more complete separation the sexes our plant would advantageous the principle the division labour individuals with this tendency more and more increased would continually favoured selected until last complete separation the sexes would effected let now turn the nectar feeding insects our imaginary case may suppose the plant which have been slowly increasing the nectar continued selection common plant and that certain insects depended main part its nectar for food could give many facts showing how anxious bees are save time for instance their habit cutting holes and sucking the nectar the bases certain flowers which they can with very little more trouble enter the mouth bearing such facts mind can see reason doubt that accidental deviation the size and form the body the curvature and length the proboscis etc far too slight appreciated might profit bee other insect that individual characterised would able obtain its food more quickly and have better chance living and leaving descendants its descendants would probably inherit tendency similar slight deviation structure the tubes the corollas the common red and incarnate clovers trifolium pratense and incarnatum not hasty glance appear differ length yet the hive bee can easily suck the nectar out the incarnate clover but not out the common red clover which visited humble bees alone that whole fields the red clover offer vain abundant supply precious nectar the hive bee thus might great advantage the hive bee have slightly longer differently constructed proboscis the other hand have found experiment that the fertility clover greatly depends bees visiting and moving parts the corolla push the pollen the stigmatic surface hence again humble bees were become rare any country might great advantage the red clover have shorter more deeply divided tube its corolla that the hive bee could visit its flowers thus can understand how flower and bee might slowly become either simultaneously one after the other modified and adapted the most perfect manner each other the continued preservation individuals presenting mutual and slightly favourable deviations structure well aware that this doctrine natural selection exemplified the above imaginary instances open the same objections which were first urged against sir charles lyell noble views the modern changes the earth illustrative geology but now very seldom hear the action for instance the coast waves called trifling and insignificant cause when applied the excavation gigantic valleys the formation the longest lines inland cliffs natural selection can act only the preservation and accumulation infinitesimally small inherited modifications each profitable the preserved being and modern geology has almost banished such views the excavation great valley single diluvial wave will natural selection true principle banish the belief the continued creation new organic beings any great and sudden modification their structure the intercrossing individuals must here introduce short digression the case animals and plants with separated sexes course obvious that two individuals must always unite for each birth but the case hermaphrodites this far from obvious nevertheless strongly inclined believe that with all hermaphrodites two individuals either occasionally habitually concur for the reproduction their kind this view may add was first suggested andrew knight shall presently see its importance but must here treat the subject with extreme brevity though have the materials prepared for ample discussion all vertebrate animals all insects and some other large groups animals pair for each birth modern research has much diminished the number supposed hermaphrodites and real hermaphrodites large number pair that two individuals regularly unite for reproduction which all that concerns but still there are many hermaphrodite animals which certainly not habitually pair and vast majority plants are hermaphrodites what reason may asked there for supposing these cases that two individuals ever concur reproduction impossible here enter details must trust some general considerations alone the first place have collected large body facts showing accordance with the almost universal belief breeders that with animals and plants cross between different varieties between individuals the same variety but another strain gives vigour and fertility the offspring and the other hand that close interbreeding diminishes vigour and fertility that these facts alone incline believe that general law nature utterly ignorant though the meaning the law that organic being self fertilises itself for eternity generations but that cross with another individual occasionally perhaps very long intervals indispensable the belief that this law nature can think understand several large classes facts such the following which any other view are inexplicable every hybridizer knows how unfavourable exposure wet the fertilisation flower yet what multitude flowers have their anthers and stigmas fully exposed the weather but occasional cross indispensable the fullest freedom for the entrance pollen from another individual will explain this state exposure more especially the plant own anthers and pistil generally stand close together that self fertilisation seems almost inevitable many flowers the other hand have their organs fructification closely enclosed the great papilionaceous pea family but several perhaps all such flowers there very curious adaptation between the structure the flower and the manner which bees suck the nectar for doing this they either push the flower own pollen the stigma bring pollen from another flower necessary are the visits bees papilionaceous flowers that have found experiments published elsewhere that their fertility greatly diminished these visits prevented now scarcely possible that bees should fly from flower flower and not carry pollen from one the other the great good believe the plant bees will act like camel hair pencil and quite sufficient just touch the anthers one flower and then the stigma another with the same brush ensure fertilisation but must not supposed that bees would thus produce multitude hybrids between distinct species for you bring the same brush plant own pollen and pollen from another species the former will have such prepotent effect that will invariably and completely destroy has been shown gärtner any influence from the foreign pollen when the stamens flower suddenly spring towards the pistil slowly move one after the other towards the contrivance seems adapted solely ensure self fertilisation and doubt useful for this end but the agency insects often required cause the stamens spring forward kölreuter has shown the case with the barberry and curiously this very genus which seems have special contrivance for self fertilisation well known that very closely allied forms varieties are planted near each other hardly possible raise pure seedlings largely they naturally cross many other cases far from there being any aids for self fertilisation there are special contrivances could show from the writings sprengel and from own observations which effectually prevent the stigma receiving pollen from its own flower for instance lobelia fulgens there really beautiful and elaborate contrivance which every one the infinitely numerous pollen granules are swept out the conjoined anthers each flower before the stigma that individual flower ready receive them and this flower never visited least garden insects never sets seed though placing pollen from one flower the stigma another raised plenty seedlings and whilst another species lobelia growing close which visited bees seeds freely very many other cases though there special mechanical contrivance prevent the stigma flower receiving its own pollen yet sprengel has shown and can confirm either the anthers burst before the stigma ready for fertilisation the stigma ready before the pollen that flower ready that these plants have fact separated sexes and must habitually crossed how strange are these facts how strange that the pollen and stigmatic surface the same flower though placed close together for the very purpose self fertilisation should many cases mutually useless each other how simply are these facts explained the view occasional cross with distinct individual being advantageous indispensable several varieties the cabbage radish onion and some other plants allowed seed near each other large majority have found the seedlings thus raised will turn out mongrels for instance raised 233 seedling cabbages from some plants different varieties growing near each other and these only were true their kind and some even these were not perfectly true yet the pistil each cabbage flower surrounded not only its own six stamens but those the many other flowers the same plant how then comes that such vast number the seedlings are mongrelized suspect that must arise from the pollen distinct variety having prepotent effect over flower own pollen and that this part the general law good being derived from the intercrossing distinct individuals the same species when distinct species are crossed the case directly the reverse for plant own pollen always prepotent over foreign pollen but this subject shall return future chapter the case gigantic tree covered with innumerable flowers may objected that pollen could seldom carried from tree tree and most only from flower flower the same tree and that flowers the same tree can considered distinct individuals only limited sense believe this objection valid but that nature has largely provided against giving trees strong tendency bear flowers with separated sexes when the sexes are separated although the male and female flowers may produced the same tree can see that pollen must regularly carried from flower flower and this will give better chance pollen being occasionally carried from tree tree that trees belonging all orders have their sexes more often separated than other plants find the case this country and request hooker tabulated the trees new zealand and asa gray those the united states and the result was anticipated the other hand hooker has recently informed that finds that the rule does not hold australia and have made these few remarks the sexes trees simply call attention the subject turning for very brief space animals the land there are some hermaphrodites land mollusca and earth worms but these all pair yet have not found single case terrestrial animal which fertilises itself can understand this remarkable fact which offers strong contrast with terrestrial plants the view occasional cross being indispensable considering the medium which terrestrial animals live and the nature the fertilising element for know means analogous the action insects and the wind the case plants which occasional cross could effected with terrestrial animals without the concurrence two individuals aquatic animals there are many self fertilising hermaphrodites but here currents the water offer obvious means for occasional cross and the case flowers have yet failed after consultation with one the highest authorities namely professor huxley discover single case hermaphrodite animal with the organs reproduction perfectly enclosed within the body that access from without and the occasional influence distinct individual can shown physically impossible cirripedes long appeared present case very great difficulty under this point view but have been enabled fortunate chance elsewhere prove that two individuals though both are self fertilising hermaphrodites sometimes cross must have struck most naturalists strange anomaly that the case both animals and plants species the same family and even the same genus though agreeing closely with each other almost their whole organisation yet are not rarely some them hermaphrodites and some them unisexual but fact all hermaphrodites occasionally intercross with other individuals the difference between hermaphrodites and unisexual species far function concerned becomes very small from these several considerations and from the many special facts which have collected but which not here able give strongly inclined suspect that both the vegetable and animal kingdoms occasional intercross with distinct individual law nature well aware that there are this view many cases difficulty some which trying investigate finally then may conclude that many organic beings cross between two individuals obvious necessity for each birth many others occurs perhaps only long intervals but none suspect can self fertilisation for perpetuity circumstances favourable natural selection this extremely intricate subject large amount inheritable and diversified variability favourable but believe mere individual differences suffice for the work large number individuals giving better chance for the appearance within any given period profitable variations will compensate for lesser amount variability each individual and believe extremely important element success though nature grants vast periods time for the work natural selection she does not grant indefinite period for all organic beings are striving may said seize each place the economy nature any one species does not become modified and improved corresponding degree with its competitors will soon exterminated man methodical selection breeder selects for some definite object and free intercrossing will wholly stop his work but when many men without intending alter the breed have nearly common standard perfection and all try get and breed from the best animals much improvement and modification surely but slowly follow from this unconscious process selection notwithstanding large amount crossing with inferior animals thus will nature for within confined area with some place its polity not perfectly occupied might natural selection will always tend preserve all the individuals varying the right direction though different degrees better fill the unoccupied place but the area large its several districts will almost certainly present different conditions life and then natural selection modifying and improving species the several districts there will intercrossing with the other individuals the same species the confines each and this case the effects intercrossing can hardly counterbalanced natural selection always tending modify all the individuals each district exactly the same manner the conditions each for continuous area the conditions will generally graduate away insensibly from one district another the intercrossing will most affect those animals which unite for each birth which wander much and which not breed very quick rate hence animals this nature for instance birds varieties will generally confined separated countries and this believe the case hermaphrodite organisms which cross only occasionally and likewise animals which unite for each birth but which wander little and which can increase very rapid rate new and improved variety might quickly formed any one spot and might there maintain itself body that whatever intercrossing took place would chiefly between the individuals the same new variety local variety when once thus formed might subsequently slowly spread other districts the above principle nurserymen always prefer getting seed from large body plants the same variety the chance intercrossing with other varieties thus lessened even the case slow breeding animals which unite for each birth must not overrate the effects intercrosses retarding natural selection for can bring considerable catalogue facts showing that within the same area varieties the same animal can long remain distinct from haunting different stations from breeding slightly different seasons from varieties the same kind preferring pair together intercrossing plays very important part nature keeping the individuals the same species the same variety true and uniform character will obviously thus act far more efficiently with those animals which unite for each birth but have already attempted show that have reason believe that occasional intercrosses take place with all animals and with all plants even these take place only long intervals convinced that the young thus produced will gain much vigour and fertility over the offspring from long continued self fertilisation that they will have better chance surviving and propagating their kind and thus the long run the influence intercrosses even rare intervals will great there exist organic beings which never intercross uniformity character can retained amongst them long their conditions life remain the same only through the principle inheritance and through natural selection destroying any which depart from the proper type but their conditions life change and they undergo modification uniformity character can given their modified offspring solely natural selection preserving the same favourable variations isolation also important element the process natural selection confined isolated area not very large the organic and inorganic conditions life will generally great degree uniform that natural selection will tend modify all the individuals varying species throughout the area the same manner relation the same conditions intercrosses also with the individuals the same species which otherwise would have inhabited the surrounding and differently circumstanced districts will prevented but isolation probably acts more efficiently checking the immigration better adapted organisms after any physical change such climate elevation the land etc and thus new places the natural economy the country are left open for the old inhabitants struggle for and become adapted through modifications their structure and constitution lastly isolation checking immigration and consequently competition will give time for any new variety slowly improved and this may sometimes importance the production new species however isolated area very small either from being surrounded barriers from having very peculiar physical conditions the total number the individuals supported will necessarily very small and fewness individuals will greatly retard the production new species through natural selection decreasing the chance the appearance favourable variations turn nature test the truth these remarks and look any small isolated area such oceanic island although the total number the species inhabiting will found small shall see our chapter geographical distribution yet these species very large proportion are endemic that have been produced there and nowhere else hence oceanic island first sight seems have been highly favourable for the production new species but may thus greatly deceive ourselves for ascertain whether small isolated area large open area like continent has been most favourable for the production new organic forms ought make the comparison within equal times and this are incapable doing although not doubt that isolation considerable importance the production new species the whole inclined believe that largeness area more importance more especially the production species which will prove capable enduring for long period and spreading widely throughout great and open area not only will there better chance favourable variations arising from the large number individuals the same species there supported but the conditions life are infinitely complex from the large number already existing species and some these many species become modified and improved others will have improved corresponding degree they will exterminated each new form also soon has been much improved will able spread over the open and continuous area and will thus come into competition with many others hence more new places will formed and the competition fill them will more severe large than small and isolated area moreover great areas though now continuous owing oscillations level will often have recently existed broken condition that the good effects isolation will generally certain extent have concurred finally conclude that although small isolated areas probably have been some respects highly favourable for the production new species yet that the course modification will generally have been more rapid large areas and what more important that the new forms produced large areas which already have been victorious over many competitors will those that will spread most widely will give rise most new varieties and species and will thus play important part the changing history the organic world can perhaps these views understand some facts which will again alluded our chapter geographical distribution for instance that the productions the smaller continent australia have formerly yielded and apparently are now yielding before those the larger europæo asiatic area thus also that continental productions have everywhere become largely naturalised islands small island the race for life will have been less severe and there will have been less modification and less extermination hence perhaps comes that the flora madeira according oswald heer resembles the extinct tertiary flora europe all fresh water basins taken together make small area compared with that the sea the land and consequently the competition between fresh water productions will have been less severe than elsewhere new forms will have been more slowly formed and old forms more slowly exterminated and fresh water that find seven genera ganoid fishes remnants once preponderant order and fresh water find some the most anomalous forms now known the world the ornithorhynchus and lepidosiren which like fossils connect certain extent orders now widely separated the natural scale these anomalous forms may almost called living fossils they have endured the present day from having inhabited confined area and from having thus been exposed less severe competition sum the circumstances favourable and unfavourable natural selection far the extreme intricacy the subject permits conclude looking the future that for terrestrial productions large continental area which will probably undergo many oscillations level and which consequently will exist for long periods broken condition will the most favourable for the production many new forms life likely endure long and spread widely for the area will first have existed continent and the inhabitants this period numerous individuals and kinds will have been subjected very severe competition when converted subsidence into large separate islands there will still exist many individuals the same species each island intercrossing the confines the range each species will thus checked after physical changes any kind immigration will prevented that new places the polity each island will have filled modifications the old inhabitants and time will allowed for the varieties each become well modified and perfected when renewed elevation the islands shall converted into continental area there will again severe competition the most favoured improved varieties will enabled spread there will much extinction the less improved forms and the relative proportional numbers the various inhabitants the renewed continent will again changed and again there will fair field for natural selection improve still further the inhabitants and thus produce new species that natural selection will always act with extreme slowness fully admit its action depends there being places the polity nature which can better occupied some the inhabitants the country undergoing modification some kind the existence such places will often depend physical changes which are generally very slow and the immigration better adapted forms having been checked but the action natural selection will probably still oftener depend some the inhabitants becoming slowly modified the mutual relations many the other inhabitants being thus disturbed nothing can effected unless favourable variations occur and variation itself apparently always very slow process the process will often greatly retarded free intercrossing many will exclaim that these several causes are amply sufficient wholly stop the action natural selection not believe the other hand believe that natural selection will always act very slowly often only long intervals time and generally only very few the inhabitants the same region the same time further believe that this very slow intermittent action natural selection accords perfectly well with what geology tells the rate and manner which the inhabitants this world have changed slow though the process selection may feeble man can much his powers artificial selection can see limit the amount change the beauty and infinite complexity the coadaptations between all organic beings one with another and with their physical conditions life which may effected the long course time nature power selection extinction this subject will more fully discussed our chapter geology but must here alluded from being intimately connected with natural selection natural selection acts solely through the preservation variations some way advantageous which consequently endure but from the high geometrical powers increase all organic beings each area already fully stocked with inhabitants follows that each selected and favoured form increases number will the less favoured forms decrease and become rare rarity geology tells the precursor extinction can also see that any form represented few individuals will during fluctuations the seasons the number its enemies run good chance utter extinction but may further than this for new forms are continually and slowly being produced unless believe that the number specific forms goes perpetually and almost indefinitely increasing numbers inevitably must become extinct that the number specific forms has not indefinitely increased geology shows plainly and indeed can see reason why they should not have thus increased for the number places the polity nature not indefinitely great not that have any means knowing that any one region has yet got its maximum species probably region yet fully stocked for the cape good hope where more species plants are crowded together than any other quarter the world some foreign plants have become naturalised without causing far know the extinction any natives furthermore the species which are most numerous individuals will have the best chance producing within any given period favourable variations have evidence this the facts given the second chapter showing that the common species which afford the greatest number recorded varieties incipient species hence rare species will less quickly modified improved within any given period and they will consequently beaten the race for life the modified descendants the commoner species from these several considerations think inevitably follows that new species the course time are formed through natural selection others will become rarer and rarer and finally extinct the forms which stand closest competition with those undergoing modification and improvement will naturally suffer most and have seen the chapter the struggle for existence that the most closely allied forms varieties the same species and species the same genus related genera which from having nearly the same structure constitution and habits generally come into the severest competition with each other consequently each new variety species during the progress its formation will generally press hardest its nearest kindred and tend exterminate them see the same process extermination amongst our domesticated productions through the selection improved forms man many curious instances could given showing how quickly new breeds cattle sheep and other animals and varieties flowers take the place older and inferior kinds yorkshire historically known that the ancient black cattle were displaced the long horns and that these were swept away the short horns quote the words agricultural writer some murderous pestilence divergence character the principle which have designated this term high importance theory and explains believe several important facts the first place varieties even strongly marked ones though having somewhat the character species shown the hopeless doubts many cases how rank them yet certainly differ from each other far less than good and distinct species nevertheless according view varieties are species the process formation are have called them incipient species how then does the lesser difference between varieties become augmented into the greater difference between species that this does habitually happen must infer from most the innumerable species throughout nature presenting well marked differences whereas varieties the supposed prototypes and parents future well marked species present slight and ill defined differences mere chance may call might cause one variety differ some character from its parents and the offspring this variety again differ from its parent the very same character and greater degree but this alone would never account for habitual and large amount difference that between varieties the same species and species the same genus has always been practice let seek light this head from our domestic productions shall here find something analogous fancier struck pigeon having slightly shorter beak another fancier struck pigeon having rather longer beak and the acknowledged principle that fanciers not and will not admire medium standard but like extremes they both has actually occurred with tumbler pigeons choosing and breeding from birds with longer and longer beaks with shorter and shorter beaks again may suppose that early period one man preferred swifter horses another stronger and more bulky horses the early differences would very slight the course time from the continued selection swifter horses some breeders and stronger ones others the differences would become greater and would noted forming two sub breeds finally after the lapse centuries the sub breeds would become converted into two well established and distinct breeds the differences slowly become greater the inferior animals with intermediate characters being neither very swift nor very strong will have been neglected and will have tended disappear here then see man productions the action what may called the principle divergence causing differences first barely appreciable steadily increase and the breeds diverge character both from each other and from their common parent but how may asked can any analogous principle apply nature believe can and does apply most efficiently from the simple circumstance that the more diversified the descendants from any one species become structure constitution and habits much will they better enabled seize many and widely diversified places the polity nature and enabled increase numbers can clearly see this the case animals with simple habits take the case carnivorous quadruped which the number that can supported any country has long ago arrived its full average its natural powers increase allowed act can succeed increasing the country not undergoing any change its conditions only its varying descendants seizing places present occupied other animals some them for instance being enabled feed new kinds prey either dead alive some inhabiting new stations climbing trees frequenting water and some perhaps becoming less carnivorous the more diversified habits and structure the descendants our carnivorous animal became the more places they would enabled occupy what applies one animal will apply throughout all time all animals that they vary for otherwise natural selection can nothing will with plants has been experimentally proved that plot ground sown with one species grass and similar plot sown with several distinct genera grasses greater number plants and greater weight dry herbage can thus raised the same has been found hold good when first one variety and then several mixed varieties wheat have been sown equal spaces ground hence any one species grass were varying and those varieties were continually selected which differed from each other all the same manner distinct species and genera grasses differ from each other greater number individual plants this species grass including its modified descendants would succeed living the same piece ground and well know that each species and each variety grass annually sowing almost countless seeds and thus may said striving its utmost increase its numbers consequently cannot doubt that the course many thousands generations the most distinct varieties any one species grass would always have the best chance succeeding and increasing numbers and thus supplanting the less distinct varieties and varieties when rendered very distinct from each other take the rank species the truth the principle that the greatest amount life can supported great diversification structure seen under many natural circumstances extremely small area especially freely open immigration and where the contest between individual and individual must severe always find great diversity its inhabitants for instance found that piece turf three feet four size which had been exposed for many years exactly the same conditions supported twenty species plants and these belonged eighteen genera and eight orders which shows how much these plants differed from each other with the plants and insects small and uniform islets and small ponds fresh water farmers find that they can raise most food rotation plants belonging the most different orders nature follows what may called simultaneous rotation most the animals and plants which live close round any small piece ground could live supposing not any way peculiar its nature and may said striving the utmost live there but seen that where they come into the closest competition with each other the advantages diversification structure with the accompanying differences habit and constitution determine that the inhabitants which thus jostle each other most closely shall general rule belong what call different genera and orders the same principle seen the naturalisation plants through man agency foreign lands might have been expected that the plants which have succeeded becoming naturalised any land would generally have been closely allied the indigenes for these are commonly looked specially created and adapted for their own country might also perhaps have been expected that naturalised plants would have belonged few groups more especially adapted certain stations their new homes but the case very different and alph candolle has well remarked his great and admirable work that floras gain naturalisation proportionally with the number the native genera and species far more new genera than new species give single instance the last edition asa gray manual the flora the northern united states 260 naturalised plants are enumerated and these belong 162 genera thus see that these naturalised plants are highly diversified nature they differ moreover large extent from the indigenes for out the 162 genera less than 100 genera are not there indigenous and thus large proportional addition made the genera these states considering the nature the plants animals which have struggled successfully with the indigenes any country and have there become naturalised can gain some crude idea what manner some the natives would have had modified order have gained advantage over the other natives and may think least safely infer that diversification structure amounting new generic differences would have been profitable them the advantage diversification the inhabitants the same region fact the same that the physiological division labour the organs the same individual body subject well elucidated milne edwards physiologist doubts that stomach being adapted digest vegetable matter alone flesh alone draws most nutriment from these substances the general economy any land the more widely and perfectly the animals and plants are diversified for different habits life will greater number individuals capable there supporting themselves set animals with their organisation but little diversified could hardly compete with set more perfectly diversified structure may doubted for instance whether the australian marsupials which are divided into groups differing but little from each other and feebly representing waterhouse and others have remarked our carnivorous ruminant and rodent mammals could successfully compete with these well pronounced orders the australian mammals see the process diversification early and incomplete stage development after the foregoing discussion which ought have been much amplified may think assume that the modified descendants any one species will succeed much the better they become more diversified structure and are thus enabled encroach places occupied other beings now let see how this principle great benefit being derived from divergence character combined with the principles natural selection and extinction will tend act the accompanying diagram will aid understanding this rather perplexing subject let represent the species genus large its own country these species are supposed resemble each other unequal degrees generally the case nature and represented the diagram the letters standing unequal distances have said large genus because have seen the second chapter that average more the species large genera vary than small genera and the varying species the large genera present greater number varieties have also seen that the species which are the commonest and the most widely diffused vary more than rare species with restricted ranges let common widely diffused and varying species belonging genus large its own country the little fan diverging dotted lines unequal lengths proceeding from may represent its varying offspring the variations are supposed extremely slight but the most diversified nature they are not supposed all appear simultaneously but often after long intervals time nor are they all supposed endure for equal periods only those variations which are some way profitable will preserved naturally selected and here the importance the principle benefit being derived from divergence character comes for this will generally lead the most different divergent variations represented the outer dotted lines being preserved and accumulated natural selection when dotted line reaches one the horizontal lines and there marked small numbered letter sufficient amount variation supposed have been accumulated have formed fairly well marked variety such would thought worthy record systematic work the intervals between the horizontal lines the diagram may represent each thousand generations but would have been better each had represented ten thousand generations after thousand generations species supposed have produced two fairly well marked varieties namely and these two varieties will generally continue exposed the same conditions which made their parents variable and the tendency variability itself hereditary consequently they will tend vary and generally vary nearly the same manner their parents varied moreover these two varieties being only slightly modified forms will tend inherit those advantages which made their common parent more numerous than most the other inhabitants the same country they will likewise partake those more general advantages which made the genus which the parent species belonged large genus its own country and these circumstances know favourable the production new varieties then these two varieties variable the most divergent their variations will generally preserved during the next thousand generations and after this interval variety supposed the diagram have produced variety which will owing the principle divergence differ more from than did variety variety supposed have produced two varieties namely and differing from each other and more considerably from their common parent may continue the process similar steps for any length time some the varieties after each thousand generations producing only single variety but more and more modified condition some producing two three varieties and some failing produce any thus the varieties modified descendants proceeding from the common parent will generally increasing number and diverging character the diagram the process represented the ten thousandth generation and under condensed and simplified form the fourteen thousandth generation but must here remark that not suppose that the process ever goes regularly represented the diagram though itself made somewhat irregular far from thinking that the most divergent varieties will invariably prevail and multiply medium form may often long endure and may may not produce more than one modified descendant for natural selection will always act according the nature the places which are either unoccupied not perfectly occupied other beings and this will depend infinitely complex relations but general rule the more diversified structure the descendants from any one species can rendered the more places they will enabled seize and the more their modified progeny will increased our diagram the line succession broken regular intervals small numbered letters marking the successive forms which have become sufficiently distinct recorded varieties but these breaks are imaginary and might have been inserted anywhere after intervals long enough have allowed the accumulation considerable amount divergent variation all the modified descendants from common and widely diffused species belonging large genus will tend partake the same advantages which made their parent successful life they will generally multiplying number well diverging character this represented the diagram the several divergent branches proceeding from the modified offspring from the later and more highly improved branches the lines descent will probable often take the place and destroy the earlier and less improved branches this represented the diagram some the lower branches not reaching the upper horizontal lines some cases not doubt that the process modification will confined single line descent and the number the descendants will not increased although the amount divergent modification may have been increased the successive generations this case would represented the diagram all the lines proceeding from were removed excepting that from the same way for instance the english race horse and english pointer have apparently both gone slowly diverging character from their original stocks without either having given off any fresh branches races after ten thousand generations species supposed have produced three forms and which from having diverged character during the successive generations will have come differ largely but perhaps unequally from each other and from their common parent suppose the amount change between each horizontal line our diagram excessively small these three forms may still only well marked varieties they may have arrived the doubtful category sub species but have only suppose the steps the process modification more numerous greater amount convert these three forms into well defined species thus the diagram illustrates the steps which the small differences distinguishing varieties are increased into the larger differences distinguishing species continuing the same process for greater number generations shown the diagram condensed and simplified manner get eight species marked the letters between and all descended from thus believe species are multiplied and genera are formed large genus probable that more than one species would vary the diagram have assumed that second species has produced analogous steps after ten thousand generations either two well marked varieties and two species according the amount change supposed represented between the horizontal lines after fourteen thousand generations six new species marked the letters are supposed have been produced each genus the species which are already extremely different character will generally tend produce the greatest number modified descendants for these will have the best chance filling new and widely different places the polity nature hence the diagram have chosen the extreme species and the nearly extreme species those which have largely varied and have given rise new varieties and species the other nine species marked capital letters our original genus may for long period continue transmitting unaltered descendants and this shown the diagram the dotted lines not prolonged far upwards from want space but during the process modification represented the diagram another our principles namely that extinction will have played important part each fully stocked country natural selection necessarily acts the selected form having some advantage the struggle for life over other forms there will constant tendency the improved descendants any one species supplant and exterminate each stage descent their predecessors and their original parent for should remembered that the competition will generally most severe between those forms which are most nearly related each other habits constitution and structure hence all the intermediate forms between the earlier and later states that between the less and more improved state species well the original parent species itself will generally tend become extinct probably will with many whole collateral lines descent which will conquered later and improved lines descent however the modified offspring species get into some distinct country become quickly adapted some quite new station which child and parent not come into competition both may continue exist then our diagram assumed represent considerable amount modification species and all the earlier varieties will have become extinct having been replaced eight new species and will have been replaced six new species but may further than this the original species our genus were supposed resemble each other unequal degrees generally the case nature species being more nearly related and than the other species and species more than the others these two species and were also supposed very common and widely diffused species that they must originally have had some advantage over most the other species the genus their modified descendants fourteen number the fourteen thousandth generation will probably have inherited some the same advantages they have also been modified and improved diversified manner each stage descent have become adapted many related places the natural economy their country seems therefore extremely probable that they will have taken the places and thus exterminated not only their parents and but likewise some the original species which were most nearly related their parents hence very few the original species will have transmitted offspring the fourteen thousandth generation may suppose that only one the two species which were least closely related the other nine original species has transmitted descendants this late stage descent the new species our diagram descended from the original eleven species will now fifteen number owing the divergent tendency natural selection the extreme amount difference character between species and will much greater than that between the most different the original eleven species the new species moreover will allied each other widely different manner the eight descendants from the three marked will nearly related from having recently branched off from and from having diverged earlier period from will some degree distinct from the three first named species and lastly and will nearly related one the other but from having diverged the first commencement the process modification will widely different from the other five species and may constitute sub genus even distinct genus the six descendants from will form two sub genera even genera but the original species differed largely from standing nearly the extreme points the original genus the six descendants from will owing inheritance differ considerably from the eight descendants from the two groups moreover are supposed have gone diverging different directions the intermediate species also and this very important consideration which connected the original species and have all become excepting extinct and have left descendants hence the six new species descended from and the eight descended from will have ranked very distinct genera even distinct sub families thus believe that two more genera are produced descent with modification from two more species the same genus and the two more parent species are supposed have descended from some one species earlier genus our diagram this indicated the broken lines beneath the capital letters converging sub branches downwards towards single point this point representing single species the supposed single parent our several new sub genera and genera worth while reflect for moment the character the new species f14 which supposed not have diverged much character but have retained the form either unaltered altered only slight degree this case its affinities the other fourteen new species will curious and circuitous nature having descended from form which stood between the two parent species and now supposed extinct and unknown will some degree intermediate character between the two groups descended from these species but these two groups have gone diverging character from the type their parents the new species f14 will not directly intermediate between them but rather between types the two groups and every naturalist will able bring some such case before his mind the diagram each horizontal line has hitherto been supposed represent thousand generations but each may represent million hundred million generations and likewise section the successive strata the earth crust including extinct remains shall when come our chapter geology have refer again this subject and think shall then see that the diagram throws light the affinities extinct beings which though generally belonging the same orders families genera with those now living yet are often some degree intermediate character between existing groups and can understand this fact for the extinct species lived very ancient epochs when the branching lines descent had diverged less see reason limit the process modification now explained the formation genera alone our diagram suppose the amount change represented each successive group diverging dotted lines very great the forms marked those marked and and those marked will form three very distinct genera shall also have two very distinct genera descended from and these latter two genera both from continued divergence character and from inheritance from different parent will differ widely from the three genera descended from the two little groups genera will form two distinct families even orders according the amount divergent modification supposed represented the diagram and the two new families orders will have descended from two species the original genus and these two species are supposed have descended from one species still more ancient and unknown genus have seen that each country the species the larger genera which oftenest present varieties incipient species this indeed might have been expected for natural selection acts through one form having some advantage over other forms the struggle for existence will chiefly act those which already have some advantage and the largeness any group shows that its species have inherited from common ancestor some advantage common hence the struggle for the production new and modified descendants will mainly lie between the larger groups which are all trying increase number one large group will slowly conquer another large group reduce its numbers and thus lessen its chance further variation and improvement within the same large group the later and more highly perfected sub groups from branching out and seizing many new places the polity nature will constantly tend supplant and destroy the earlier and less improved sub groups small and broken groups and sub groups will finally tend disappear looking the future can predict that the groups organic beings which are now large and triumphant and which are least broken that which yet have suffered least extinction will for long period continue increase but which groups will ultimately prevail man can predict for well know that many groups formerly most extensively developed have now become extinct looking still more remotely the future may predict that owing the continued and steady increase the larger groups multitude smaller groups will become utterly extinct and leave modified descendants and consequently that the species living any one period extremely few will transmit descendants remote futurity shall have return this subject the chapter classification but may add that this view extremely few the more ancient species having transmitted descendants and the view all the descendants the same species making class can understand how that there exist but very few classes each main division the animal and vegetable kingdoms although extremely few the most ancient species may now have living and modified descendants yet the most remote geological period the earth may have been well peopled with many species many genera families orders and classes the present day summary the chapter during the long course ages and under varying conditions life organic beings vary all the several parts their organisation and think this cannot disputed there owing the high geometrical powers increase each species some age season year severe struggle for life and this certainly cannot disputed then considering the infinite complexity the relations all organic beings each other and their conditions existence causing infinite diversity structure constitution and habits advantageous them think would most extraordinary fact variation ever had occurred useful each being own welfare the same way many variations have occurred useful man but variations useful any organic being occur assuredly individuals thus characterised will have the best chance being preserved the struggle for life and from the strong principle inheritance they will tend produce offspring similarly characterised this principle preservation have called for the sake brevity natural selection natural selection the principle qualities being inherited corresponding ages can modify the egg seed young easily the adult amongst many animals sexual selection will give its aid ordinary selection assuring the most vigorous and best adapted males the greatest number offspring sexual selection will also give characters useful the males alone their struggles with other males whether natural selection has really thus acted nature modifying and adapting the various forms life their several conditions and stations must judged the general tenour and balance evidence given the following chapters but already see how entails extinction and how largely extinction has acted the world history geology plainly declares natural selection also leads divergence character for more living beings can supported the same area the more they diverge structure habits and constitution which see proof looking the inhabitants any small spot naturalised productions therefore during the modification the descendants any one species and during the incessant struggle all species increase numbers the more diversified these descendants become the better will their chance succeeding the battle life thus the small differences distinguishing varieties the same species will steadily tend increase till they come equal the greater differences between species the same genus even distinct genera have seen that the common the widely diffused and widely ranging species belonging the larger genera which vary most and these will tend transmit their modified offspring that superiority which now makes them dominant their own countries natural selection has just been remarked leads divergence character and much extinction the less improved and intermediate forms life these principles believe the nature the affinities all organic beings may explained truly wonderful fact the wonder which are apt overlook from familiarity that all animals and all plants throughout all time and space should related each other group subordinate group the manner which everywhere behold namely varieties the same species most closely related together species the same genus less closely and unequally related together forming sections and sub genera species distinct genera much less closely related and genera related different degrees forming sub families families orders sub classes and classes the several subordinate groups any class cannot ranked single file but seem rather clustered round points and these round other points and almost endless cycles the view that each species has been independently created can see explanation this great fact the classification all organic beings but the best judgment explained through inheritance and the complex action natural selection entailing extinction and divergence character have seen illustrated the diagram the affinities all the beings the same class have sometimes been represented great tree believe this simile largely speaks the truth the green and budding twigs may represent existing species and those produced during each former year may represent the long succession extinct species each period growth all the growing twigs have tried branch out all sides and overtop and kill the surrounding twigs and branches the same manner species and groups species have tried overmaster other species the great battle for life the limbs divided into great branches and these into lesser and lesser branches were themselves once when the tree was small budding twigs and this connexion the former and present buds ramifying branches may well represent the classification all extinct and living species groups subordinate groups the many twigs which flourished when the tree was mere bush only two three now grown into great branches yet survive and bear all the other branches with the species which lived during long past geological periods very few now have living and modified descendants from the first growth the tree many limb and branch has decayed and dropped off and these lost branches various sizes may represent those whole orders families and genera which have now living representatives and which are known only from having been found fossil state here and there see thin straggling branch springing from fork low down tree and which some chance has been favoured and still alive its summit occasionally see animal like the ornithorhynchus lepidosiren which some small degree connects its affinities two large branches life and which has apparently been saved from fatal competition having inhabited protected station buds give rise growth fresh buds and these vigorous branch out and overtop all sides many feebler branch generation believe has been with the great tree life which fills with its dead and broken branches the crust the earth and covers the surface with its ever branching and beautiful ramifications chapter laws variation effects external conditions use and disuse combined with natural selection organs flight and vision acclimatisation correlation growth compensation and economy growth false correlations multiple rudimentary and lowly organised structures variable parts developed unusual manner are highly variable specific characters more variable than generic secondary sexual characters variable species the same genus vary analogous manner reversions long lost characters summary have hitherto sometimes spoken the variations common and multiform organic beings under domestication and lesser degree those state nature had been due chance this course wholly incorrect expression but serves acknowledge plainly our ignorance the cause each particular variation some authors believe much the function the reproductive system produce individual differences very slight deviations structure make the child like its parents but the much greater variability well the greater frequency monstrosities under domestication cultivation than under nature leads believe that deviations structure are some way due the nature the conditions life which the parents and their more remote ancestors have been exposed during several generations have remarked the first chapter but long catalogue facts which cannot here given would necessary show the truth the remark that the reproductive system eminently susceptible changes the conditions life and this system being functionally disturbed the parents chiefly attribute the varying plastic condition the offspring the male and female sexual elements seem affected before that union takes place which form new being the case sporting plants the bud which its earliest condition does not apparently differ essentially from ovule alone affected but why because the reproductive system disturbed this that part should vary more less are profoundly ignorant nevertheless can here and there dimly catch faint ray light and may feel sure that there must some cause for each deviation structure however slight how much direct effect difference climate food etc produces any being extremely doubtful impression that the effect extremely small the case animals but perhaps rather more that plants may least safely conclude that such influences cannot have produced the many striking and complex adaptations structure between one organic being and another which see everywhere throughout nature some little influence may attributed climate food etc thus forbes speaks confidently that shells their southern limit and when living shallow water are more brightly coloured than those the same species further north from greater depths gould believes that birds the same species are more brightly coloured under clear atmosphere than when living islands near the coast with insects wollaston convinced that residence near the sea affects their colours moquin tandon gives list plants which when growing near the sea shore have their leaves some degree fleshy though not elsewhere fleshy several other such cases could given the fact varieties one species when they range into the zone habitation other species often acquiring very slight degree some the characters such species accords with our view that species all kinds are only well marked and permanent varieties thus the species shells which are confined tropical and shallow seas are generally brighter coloured than those confined cold and deeper seas the birds which are confined continents are according gould brighter coloured than those islands the insect species confined sea coasts every collector knows are often brassy lurid plants which live exclusively the sea side are very apt have fleshy leaves who believes the creation each species will have say that this shell for instance was created with bright colours for warm sea but that this other shell became bright coloured variation when ranged into warmer shallower waters when variation the slightest use being cannot tell how much attribute the accumulative action natural selection and how much the conditions life thus well known furriers that animals the same species have thicker and better fur the more severe the climate under which they have lived but who can tell how much this difference may due the warmest clad individuals having been favoured and preserved during many generations and how much the direct action the severe climate for would appear that climate has some direct action the hair our domestic quadrupeds instances could given the same variety being produced under conditions life different can well conceived and the other hand different varieties being produced from the same species under the same conditions such facts show how indirectly the conditions life must act again innumerable instances are known every naturalist species keeping true not varying all although living under the most opposite climates such considerations these incline lay very little weight the direct action the conditions life indirectly already remarked they seem play important part affecting the reproductive system and thus inducing variability and natural selection will then accumulate all profitable variations however slight until they become plainly developed and appreciable effects use and disuse from the facts alluded the first chapter think there can little doubt that use our domestic animals strengthens and enlarges certain parts and disuse diminishes them and that such modifications are inherited under free nature can have standard comparison which judge the effects long continued use disuse for know not the parent forms but many animals have structures which can explained the effects disuse professor owen has remarked there greater anomaly nature than bird that cannot fly yet there are several this state the logger headed duck south america can only flap along the surface the water and has its wings nearly the same condition the domestic aylesbury duck the larger ground feeding birds seldom take flight except escape danger believe that the nearly wingless condition several birds which now inhabit have lately inhabited several oceanic islands tenanted beast prey has been caused disuse the ostrich indeed inhabits continents and exposed danger from which cannot escape flight but kicking can defend itself from enemies well any the smaller quadrupeds may imagine that the early progenitor the ostrich had habits like those bustard and that natural selection increased successive generations the size and weight its body its legs were used more and its wings less until they became incapable flight kirby has remarked and have observed the same fact that the anterior tarsi feet many male dung feeding beetles are very often broken off examined seventeen specimens his own collection and not one had even relic left the onites apelles the tarsi are habitually lost that the insect has been described not having them some other genera they are present but rudimentary condition the ateuchus sacred beetle the egyptians they are totally deficient there not sufficient evidence induce believe that mutilations are ever inherited and should prefer explaining the entire absence the anterior tarsi ateuchus and their rudimentary condition some other genera the long continued effects disuse their progenitors for the tarsi are almost always lost many dung feeding beetles they must lost early life and therefore cannot much used these insects some cases might easily put down disuse modifications structure which are wholly mainly due natural selection wollaston has discovered the remarkable fact that 200 beetles out the 550 species inhabiting madeira are far deficient wings that they cannot fly and that the twenty nine endemic genera less than twenty three genera have all their species this condition several facts namely that beetles many parts the world are very frequently blown sea and perish that the beetles madeira observed wollaston lie much concealed until the wind lulls and the sun shines that the proportion wingless beetles larger the exposed dezertas than madeira itself and especially the extraordinary fact strongly insisted wollaston the almost entire absence certain large groups beetles elsewhere excessively numerous and which groups have habits life almost necessitating frequent flight these several considerations have made believe that the wingless condition many madeira beetles mainly due the action natural selection but combined probably with disuse for during thousands successive generations each individual beetle which flew least either from its wings having been ever little less perfectly developed from indolent habit will have had the best chance surviving from not being blown out sea and the other hand those beetles which most readily took flight will oftenest have been blown sea and thus have been destroyed the insects madeira which are not ground feeders and which the flower feeding coleoptera and lepidoptera must habitually use their wings gain their subsistence have wollaston suspects their wings not all reduced but even enlarged this quite compatible with the action natural selection for when new insect first arrived the island the tendency natural selection enlarge reduce the wings would depend whether greater number individuals were saved successfully battling with the winds giving the attempt and rarely never flying with mariners shipwrecked near coast would have been better for the good swimmers they had been able swim still further whereas would have been better for the bad swimmers they had not been able swim all and had stuck the wreck the eyes moles and some burrowing rodents are rudimentary size and some cases are quite covered skin and fur this state the eyes probably due gradual reduction from disuse but aided perhaps natural selection south america burrowing rodent the tuco tuco ctenomys even more subterranean its habits than the mole and was assured spaniard who had often caught them that they were frequently blind one which kept alive was certainly this condition the cause appeared dissection having been inflammation the nictitating membrane frequent inflammation the eyes must injurious any animal and eyes are certainly not indispensable animals with subterranean habits reduction their size with the adhesion the eyelids and growth fur over them might such case advantage and natural selection would constantly aid the effects disuse well known that several animals belonging the most different classes which inhabit the caves styria and kentucky are blind some the crabs the foot stalk for the eye remains though the eye gone the stand for the telescope there though the telescope with its glasses has been lost difficult imagine that eyes though useless could any way injurious animals living darkness attribute their loss wholly disuse one the blind animals namely the cave rat the eyes are immense size and professor silliman thought that regained after living some days the light some slight power vision the same manner madeira the wings some the insects have been enlarged and the wings others have been reduced natural selection aided use and disuse the case the cave rat natural selection seems have struggled with the loss light and have increased the size the eyes whereas with all the other inhabitants the caves disuse itself seems have done its work difficult imagine conditions life more similar than deep limestone caverns under nearly similar climate that the common view the blind animals having been separately created for the american and european caverns close similarity their organisation and affinities might have been expected but schiödte and others have remarked this not the case and the cave insects the two continents are not more closely allied than might have been anticipated from the general resemblance the other inhabitants north america and europe view must suppose that american animals having ordinary powers vision slowly migrated successive generations from the outer world into the deeper and deeper recesses the kentucky caves did european animals into the caves europe have some evidence this gradation habit for schiödte remarks animals not far remote from ordinary forms prepare the transition from light darkness next follow those that are constructed for twilight and last all those destined for total darkness the time that animal had reached after numberless generations the deepest recesses disuse will this view have more less perfectly obliterated its eyes and natural selection will often have effected other changes such increase the length the antennæ palpi compensation for blindness notwithstanding such modifications might expect still see the cave animals america affinities the other inhabitants that continent and those europe the inhabitants the european continent and this the case with some the american cave animals hear from professor dana and some the european cave insects are very closely allied those the surrounding country would most difficult give any rational explanation the affinities the blind cave animals the other inhabitants the two continents the ordinary view their independent creation that several the inhabitants the caves the old and new worlds should closely related might expect from the well known relationship most their other productions far from feeling any surprise that some the cave animals should very anomalous agassiz has remarked regard the blind fish the amblyopsis and the case with the blind proteus with reference the reptiles europe only surprised that more wrecks ancient life have not been preserved owing the less severe competition which the inhabitants these dark abodes will probably have been exposed acclimatisation habit hereditary with plants the period flowering the amount rain requisite for seeds germinate the time sleep etc and this leads say few words acclimatisation extremely common for species the same genus inhabit very hot and very cold countries and believe that all the species the same genus have descended from single parent this view correct acclimatisation must readily effected during long continued descent notorious that each species adapted the climate its own home species from arctic even from temperate region cannot endure tropical climate conversely again many succulent plants cannot endure damp climate but the degree adaptation species the climates under which they live often overrated may infer this from our frequent inability predict whether not imported plant will endure our climate and from the number plants and animals brought from warmer countries which here enjoy good health have reason believe that species state nature are limited their ranges the competition other organic beings quite much more than adaptation particular climates but whether not the adaptation generally very close have evidence the case some few plants their becoming certain extent naturally habituated different temperatures becoming acclimatised thus the pines and rhododendrons raised from seed collected hooker from trees growing different heights the himalaya were found this country possess different constitutional powers resisting cold thwaites informs that has observed similar facts ceylon and analogous observations have been made watson european species plants brought from the azores england regard animals several authentic cases could given species within historical times having largely extended their range from warmer cooler latitudes and conversely but not positively know that these animals were strictly adapted their native climate but all ordinary cases assume such the case nor know that they have subsequently become acclimatised their new homes believe that our domestic animals were originally chosen uncivilised man because they were useful and bred readily under confinement and not because they were subsequently found capable far extended transportation think the common and extraordinary capacity our domestic animals not only withstanding the most different climates but being perfectly fertile far severer test under them may used argument that large proportion other animals now state nature could easily brought bear widely different climates must not however push the foregoing argument too far account the probable origin some our domestic animals from several wild stocks the blood for instance tropical and arctic wolf wild dog may perhaps mingled our domestic breeds the rat and mouse cannot considered domestic animals but they have been transported man many parts the world and now have far wider range than any other rodent living free under the cold climate faroe the north and the falklands the south and many islands the torrid zones hence inclined look adaptation any special climate quality readily grafted innate wide flexibility constitution which common most animals this view the capacity enduring the most different climates man himself and his domestic animals and such facts that former species the elephant and rhinoceros were capable enduring glacial climate whereas the living species are now all tropical sub tropical their habits ought not looked anomalies but merely examples very common flexibility constitution brought under peculiar circumstances into play how much the acclimatisation species any peculiar climate due mere habit and how much the natural selection varieties having different innate constitutions and how much both means combined very obscure question that habit custom has some influence must believe both from analogy and from the incessant advice given agricultural works even the ancient encyclopædias china very cautious transposing animals from one district another for not likely that man should have succeeded selecting many breeds and sub breeds with constitutions specially fitted for their own districts the result must think due habit the other hand can see reason doubt that natural selection will continually tend preserve those individuals which are born with constitutions best adapted their native countries treatises many kinds cultivated plants certain varieties are said withstand certain climates better than others this very strikingly shown works fruit trees published the united states which certain varieties are habitually recommended for the northern and others for the southern states and most these varieties are recent origin they cannot owe their constitutional differences habit the case the jerusalem artichoke which never propagated seed and which consequently new varieties have not been produced has even been advanced for now tender ever was proving that acclimatisation cannot effected the case also the kidney bean has been often cited for similar purpose and with much greater weight but until some one will sow during score generations his kidney beans early that very large proportion are destroyed frost and then collect seed from the few survivors with care prevent accidental crosses and then again get seed from these seedlings with the same precautions the experiment cannot said have been even tried nor let supposed that differences the constitution seedling kidney beans ever appear for account has been published how much more hardy some seedlings appeared than others the whole think may conclude that habit use and disuse have some cases played considerable part the modification the constitution and the structure various organs but that the effects use and disuse have often been largely combined with and sometimes overmastered the natural selection innate differences correlation growth mean this expression that the whole organisation tied together during its growth and development that when slight variations any one part occur and are accumulated through natural selection other parts become modified this very important subject most imperfectly understood the most obvious case that modifications accumulated solely for the good the young larva will may safely concluded affect the structure the adult the same manner any malconformation affecting the early embryo seriously affects the whole organisation the adult the several parts the body which are homologous and which early embryonic period are alike seem liable vary allied manner see this the right and left sides the body varying the same manner the front and hind legs and even the jaws and limbs varying together for the lower jaw believed homologous with the limbs these tendencies not doubt may mastered more less completely natural selection thus family stags once existed with antler only one side and this had been any great use the breed might probably have been rendered permanent natural selection homologous parts has been remarked some authors tend cohere this often seen monstrous plants and nothing more common than the union homologous parts normal structures the union the petals the corolla into tube hard parts seem affect the form adjoining soft parts believed some authors that the diversity the shape the pelvis birds causes the remarkable diversity the shape their kidneys others believe that the shape the pelvis the human mother influences pressure the shape the head the child snakes according schlegel the shape the body and the manner swallowing determine the position several the most important viscera the nature the bond correlation very frequently quite obscure geoffroy hilaire has forcibly remarked that certain malconformations very frequently and that others rarely coexist without our being able assign any reason what can more singular than the relation between blue eyes and deafness cats and the tortoise shell colour with the female sex the feathered feet and skin between the outer toes pigeons and the presence more less down the young birds when first hatched with the future colour their plumage again the relation between the hair and teeth the naked turkish dog though here probably homology comes into play with respect this latter case correlation think can hardly accidental that pick out the two orders mammalia which are most abnormal their dermal coverings viz cetacea whales and edentata armadilloes scaly ant eaters etc that these are likewise the most abnormal their teeth know case better adapted show the importance the laws correlation modifying important structures independently utility and therefore natural selection than that the difference between the outer and inner flowers some compositous and umbelliferous plants every one knows the difference the ray and central florets for instance the daisy and this difference often accompanied with the abortion parts the flower but some compositous plants the seeds also differ shape and sculpture and even the ovary itself with its accessory parts differs has been described cassini these differences have been attributed some authors pressure and the shape the seeds the ray florets some compositæ countenances this idea but the case the corolla the umbelliferæ means hooker informs species with the densest heads that the inner and outer flowers most frequently differ might have been thought that the development the ray petals drawing nourishment from certain other parts the flower had caused their abortion but some compositæ there difference the seeds the outer and inner florets without any difference the corolla possibly these several differences may connected with some difference the flow nutriment towards the central and external flowers know least that irregular flowers those nearest the axis are oftenest subject peloria and become regular may add instance this and striking case correlation that have recently observed some garden pelargoniums that the central flower the truss often loses the patches darker colour the two upper petals and that when this occurs the adherent nectary quite aborted when the colour absent from only one the two upper petals the nectary only much shortened with respect the difference the corolla the central and exterior flowers head umbel not feel all sure that sprengel idea that the ray florets serve attract insects whose agency highly advantageous the fertilisation plants these two orders far fetched may first appear and advantageous natural selection may have come into play but regard the differences both the internal and external structure the seeds which are not always correlated with any differences the flowers seems impossible that they can any way advantageous the plant yet the umbelliferæ these differences are such apparent importance the seeds being some cases according tausch orthospermous the exterior flowers and coelospermous the central flowers that the elder candolle founded his main divisions the order analogous differences hence see that modifications structure viewed systematists high value may wholly due unknown laws correlated growth and without being far can see the slightest service the species may often falsely attribute correlation growth structures which are common whole groups species and which truth are simply due inheritance for ancient progenitor may have acquired through natural selection some one modification structure and after thousands generations some other and independent modification and these two modifications having been transmitted whole group descendants with diverse habits would naturally thought correlated some necessary manner again not doubt that some apparent correlations occurring throughout whole orders are entirely due the manner alone which natural selection can act for instance alph candolle has remarked that winged seeds are never found fruits which not open should explain the rule the fact that seeds could not gradually become winged through natural selection except fruits which opened that the individual plants producing seeds which were little better fitted wafted further might get advantage over those producing seed less fitted for dispersal and this process could not possibly fruit which did not open the elder geoffroy and goethe propounded about the same period their law compensation balancement growth goethe expressed order spend one side nature forced economise the other side think this holds true certain extent with our domestic productions nourishment flows one part organ excess rarely flows least excess another part thus difficult get cow give much milk and fatten readily the same varieties the cabbage not yield abundant and nutritious foliage and copious supply oil bearing seeds when the seeds our fruits become atrophied the fruit itself gains largely size and quality our poultry large tuft feathers the head generally accompanied diminished comb and large beard diminished wattles with species state nature can hardly maintained that the law universal application but many good observers more especially botanists believe its truth will not however here give any instances for see hardly any way distinguishing between the effects the one hand part being largely developed through natural selection and another and adjoining part being reduced this same process disuse and the other hand the actual withdrawal nutriment from one part owing the excess growth another and adjoining part suspect also that some the cases compensation which have been advanced and likewise some other facts may merged under more general principle namely that natural selection continually trying economise every part the organisation under changed conditions life structure before useful becomes less useful any diminution however slight its development will seized natural selection for will profit the individual not have its nutriment wasted building useless structure can thus only understand fact with which was much struck when examining cirripedes and which many other instances could given namely that when cirripede parasitic within another and thus protected loses more less completely its own shell carapace this the case with the male ibla and truly extraordinary manner with the proteolepas for the carapace all other cirripedes consists the three highly important anterior segments the head enormously developed and furnished with great nerves and muscles but the parasitic and protected proteolepas the whole anterior part the head reduced the merest rudiment attached the bases the prehensile antennæ now the saving large and complex structure when rendered superfluous the parasitic habits the proteolepas though effected slow steps would decided advantage each successive individual the species for the struggle for life which every animal exposed each individual proteolepas would have better chance supporting itself less nutriment being wasted developing structure now become useless thus believe natural selection will always succeed the long run reducing and saving every part the organisation soon rendered superfluous without any means causing some other part largely developed corresponding degree and conversely that natural selection may perfectly well succeed largely developing any organ without requiring necessary compensation the reduction some adjoining part seems rule remarked geoffroy hilaire both varieties and species that when any part organ repeated many times the structure the same individual the vertebræ snakes and the stamens polyandrous flowers the number variable whereas the number the same part organ when occurs lesser numbers constant the same author and some botanists have further remarked that multiple parts are also very liable variation structure inasmuch this vegetative repetition use professor owen expression seems sign low organisation the foregoing remark seems connected with the very general opinion naturalists that beings low the scale nature are more variable than those which are higher presume that lowness this case means that the several parts the organisation have been but little specialised for particular functions and long the same part has perform diversified work can perhaps see why should remain variable that why natural selection should have preserved rejected each little deviation form less carefully than when the part has serve for one special purpose alone the same way that knife which has cut all sorts things may almost any shape whilst tool for some particular object had better some particular shape natural selection should never forgotten can act each part each being solely through and for its advantage rudimentary parts has been stated some authors and believe with truth are apt highly variable shall have recur the general subject rudimentary and aborted organs and will here only add that their variability seems owing their uselessness and therefore natural selection having power check deviations their structure thus rudimentary parts are left the free play the various laws growth the effects long continued disuse and the tendency reversion part developed any species extraordinary degree manner comparison with the same part allied species tends highly variable several years ago was much struck with remark nearly the above effect published waterhouse infer also from observation made professor owen with respect the length the arms the ourang outang that has come nearly similar conclusion hopeless attempt convince any one the truth this proposition without giving the long array facts which have collected and which cannot possibly here introduced can only state conviction that rule high generality aware several causes error but hope that have made due allowance for them should understood that the rule means applies any part however unusually developed unless unusually developed comparison with the same part closely allied species thus the bat wing most abnormal structure the class mammalia but the rule would not here apply because there whole group bats having wings would apply only some one species bat had its wings developed some remarkable manner comparison with the other species the same genus the rule applies very strongly the case secondary sexual characters when displayed any unusual manner the term secondary sexual characters used hunter applies characters which are attached one sex but are not directly connected with the act reproduction the rule applies males and females but females more rarely offer remarkable secondary sexual characters applies more rarely them the rule being plainly applicable the case secondary sexual characters may due the great variability these characters whether not displayed any unusual manner which fact think there can little doubt but that our rule not confined secondary sexual characters clearly shown the case hermaphrodite cirripedes and may here add that particularly attended waterhouse remark whilst investigating this order and fully convinced that the rule almost invariably holds good with cirripedes shall future work give list the more remarkable cases will here only briefly give one illustrates the rule its largest application the opercular valves sessile cirripedes rock barnacles are every sense the word very important structures and they differ extremely little even different genera but the several species one genus pyrgoma these valves present marvellous amount diversification the homologous valves the different species being sometimes wholly unlike shape and the amount variation the individuals several the species great that exaggeration state that the varieties differ more from each other the characters these important valves than other species distinct genera birds within the same country vary remarkably small degree have particularly attended them and the rule seems certainly hold good this class cannot make out that applies plants and this would seriously have shaken belief its truth had not the great variability plants made particularly difficult compare their relative degrees variability when see any part organ developed remarkable degree manner any species the fair presumption that high importance that species nevertheless the part this case eminently liable variation why should this the view that each species has been independently created with all its parts now see them can see explanation but the view that groups species have descended from other species and have been modified through natural selection think can obtain some light our domestic animals any part the whole animal neglected and selection applied that part for instance the comb the dorking fowl the whole breed will cease have nearly uniform character the breed will then said have degenerated rudimentary organs and those which have been but little specialised for any particular purpose and perhaps polymorphic groups see nearly parallel natural case for such cases natural selection either has not cannot come into full play and thus the organisation left fluctuating condition but what here more especially concerns that our domestic animals those points which the present time are undergoing rapid change continued selection are also eminently liable variation look the breeds the pigeon see what prodigious amount difference there the beak the different tumblers the beak and wattle the different carriers the carriage and tail our fantails etc these being the points now mainly attended english fanciers even the sub breeds the short faced tumbler notoriously difficult breed them nearly perfection and frequently individuals are born which depart widely from the standard there may truly said constant struggle going between the one hand the tendency reversion less modified state well innate tendency further variability all kinds and the other hand the power steady selection keep the breed true the long run selection gains the day and not expect fail far breed bird coarse common tumbler from good short faced strain but long selection rapidly going there may always expected much variability the structure undergoing modification further deserves notice that these variable characters produced man selection sometimes become attached from causes quite unknown more one sex than the other generally the male sex with the wattle carriers and the enlarged crop pouters now let turn nature when part has been developed extraordinary manner any one species compared with the other species the same genus may conclude that this part has undergone extraordinary amount modification since the period when the species branched off from the common progenitor the genus this period will seldom remote any extreme degree species very rarely endure for more than one geological period extraordinary amount modification implies unusually large and long continued amount variability which has continually been accumulated natural selection for the benefit the species but the variability the extraordinarily developed part organ has been great and long continued within period not excessively remote might general rule expect still find more variability such parts than other parts the organisation which have remained for much longer period nearly constant and this convinced the case that the struggle between natural selection the one hand and the tendency reversion and variability the other hand will the course time cease and that the most abnormally developed organs may made constant can see reason doubt hence when organ however abnormal may has been transmitted approximately the same condition many modified descendants the case the wing the bat must have existed according theory for immense period nearly the same state and thus comes more variable than any other structure only those cases which the modification has been comparatively recent and extraordinarily great that ought find the generative variability may called still present high degree for this case the variability will seldom yet have been fixed the continued selection the individuals varying the required manner and degree and the continued rejection those tending revert former and less modified condition the principle included these remarks may extended notorious that specific characters are more variable than generic explain simple example what meant some species large genus plants had blue flowers and some had red the colour would only specific character and one would surprised one the blue species varying into red conversely but all the species had blue flowers the colour would become generic character and its variation would more unusual circumstance have chosen this example because explanation not this case applicable which most naturalists would advance namely that specific characters are more variable than generic because they are taken from parts less physiological importance than those commonly used for classing genera believe this explanation partly yet only indirectly true shall however have return this subject our chapter classification would almost superfluous adduce evidence support the above statement that specific characters are more variable than generic but have repeatedly noticed works natural history that when author has remarked with surprise that some important organ part which generally very constant throughout large groups species has differed considerably closely allied species that has also been variable the individuals some the species and this fact shows that character which generally generic value when sinks value and becomes only specific value often becomes variable though its physiological importance may remain the same something the same kind applies monstrosities least geoffroy hilaire seems entertain doubt that the more organ normally differs the different species the same group the more subject individual anomalies the ordinary view each species having been independently created why should that part the structure which differs from the same part other independently created species the same genus more variable than those parts which are closely alike the several species not see that any explanation can given but the view species being only strongly marked and fixed varieties might surely expect find them still often continuing vary those parts their structure which have varied within moderately recent period and which have thus come differ state the case another manner the points which all the species genus resemble each other and which they differ from the species some other genus are called generic characters and these characters common attribute inheritance from common progenitor for can rarely have happened that natural selection will have modified several species fitted more less widely different habits exactly the same manner and these called generic characters have been inherited from remote period since that period when the species first branched off from their common progenitor and subsequently have not varied come differ any degree only slight degree not probable that they should vary the present day the other hand the points which species differ from other species the same genus are called specific characters and these specific characters have varied and come differ within the period the branching off the species from common progenitor probable that they should still often some degree variable least more variable than those parts the organisation which have for very long period remained constant connexion with the present subject will make only two other remarks think will admitted without entering details that secondary sexual characters are very variable think also will admitted that species the same group differ from each other more widely their secondary sexual characters than other parts their organisation compare for instance the amount difference between the males gallinaceous birds which secondary sexual characters are strongly displayed with the amount difference between their females and the truth this proposition will granted the cause the original variability secondary sexual characters not manifest but can see why these characters should not have been rendered constant and uniform other parts the organisation for secondary sexual characters have been accumulated sexual selection which less rigid its action than ordinary selection does not entail death but only gives fewer offspring the less favoured males whatever the cause may the variability secondary sexual characters they are highly variable sexual selection will have had wide scope for action and may thus readily have succeeded giving the species the same group greater amount difference their sexual characters than other parts their structure remarkable fact that the secondary sexual differences between the two sexes the same species are generally displayed the very same parts the organisation which the different species the same genus differ from each other this fact will give illustration two instances the first which happen stand list and the differences these cases are very unusual nature the relation can hardly accidental the same number joints the tarsi character generally common very large groups beetles but the engidæ westwood has remarked the number varies greatly and the number likewise differs the two sexes the same species again fossorial hymenoptera the manner neuration the wings character the highest importance because common large groups but certain genera the neuration differs the different species and likewise the two sexes the same species this relation has clear meaning view the subject look all the species the same genus having certainly descended from the same progenitor have the two sexes any one the species consequently whatever part the structure the common progenitor its early descendants became variable variations this part would highly probable taken advantage natural and sexual selection order fit the several species their several places the economy nature and likewise fit the two sexes the same species each other fit the males and females different habits life the males struggle with other males for the possession the females finally then conclude that the greater variability specific characters those which distinguish species from species than generic characters those which the species possess common that the frequent extreme variability any part which developed species extraordinary manner comparison with the same part its congeners and the not great degree variability part however extraordinarily may developed common whole group species that the great variability secondary sexual characters and the great amount difference these same characters between closely allied species that secondary sexual and ordinary specific differences are generally displayed the same parts the organisation are all principles closely connected together all being mainly due the species the same group having descended from common progenitor from whom they have inherited much common parts which have recently and largely varied being more likely still varying than parts which have long been inherited and have not varied natural selection having more less completely according the lapse time overmastered the tendency reversion and further variability sexual selection being less rigid than ordinary selection and variations the same parts having been accumulated natural and sexual selection and thus adapted for secondary sexual and for ordinary specific purposes distinct species present analogous variations and variety one species often assumes some the characters allied species reverts some the characters early progenitor these propositions will most readily understood looking our domestic races the most distinct breeds pigeons countries most widely apart present sub varieties with reversed feathers the head and feathers the feet characters not possessed the aboriginal rock pigeon these then are analogous variations two more distinct races the frequent presence fourteen even sixteen tail feathers the pouter may considered variation representing the normal structure another race the fantail presume that one will doubt that all such analogous variations are due the several races the pigeon having inherited from common parent the same constitution and tendency variation when acted similar unknown influences the vegetable kingdom have case analogous variation the enlarged stems roots commonly called the swedish turnip and ruta baga plants which several botanists rank varieties produced cultivation from common parent this not the case will then one analogous variation two called distinct species and these third may added namely the common turnip according the ordinary view each species having been independently created should have attribute this similarity the enlarged stems these three plants not the vera causa community descent and consequent tendency vary like manner but three separate yet closely related acts creation with pigeons however have another case namely the occasional appearance all the breeds slaty blue birds with two black bars the wings white rump bar the end the tail with the outer feathers externally edged near their bases with white all these marks are characteristic the parent rock pigeon presume that one will doubt that this case reversion and not new yet analogous variation appearing the several breeds may think confidently come this conclusion because have seen these coloured marks are eminently liable appear the crossed offspring two distinct and differently coloured breeds and this case there nothing the external conditions life cause the reappearance the slaty blue with the several marks beyond the influence the mere act crossing the laws inheritance doubt very surprising fact that characters should reappear after having been lost for many perhaps for hundreds generations but when breed has been crossed only once some other breed the offspring occasionally show tendency revert character the foreign breed for many generations some say for dozen even score generations after twelve generations the proportion blood use common expression any one ancestor only 2048 and yet see generally believed that tendency reversion retained this very small proportion foreign blood breed which has not been crossed but which both parents have lost some character which their progenitor possessed the tendency whether strong weak reproduce the lost character might was formerly remarked for all that can see the contrary transmitted for almost any number generations when character which has been lost breed reappears after great number generations the most probable hypothesis not that the offspring suddenly takes after ancestor some hundred generations distant but that each successive generation there has been tendency reproduce the character question which last under unknown favourable conditions gains ascendancy for instance probable that each generation the barb pigeon which produces most rarely blue and black barred bird there has been tendency each generation the plumage assume this colour this view hypothetical but could supported some facts and can see more abstract improbability tendency produce any character being inherited for endless number generations than quite useless rudimentary organs being all know them thus inherited indeed may sometimes observe mere tendency produce rudiment inherited for instance the common snapdragon antirrhinum rudiment fifth stamen often appears that this plant must have inherited tendency produce all the species the same genus are supposed theory have descended from common parent might expected that they would occasionally vary analogous manner that variety one species would resemble some its characters another species this other species being view only well marked and permanent variety but characters thus gained would probably unimportant nature for the presence all important characters will governed natural selection accordance with the diverse habits the species and will not left the mutual action the conditions life and similar inherited constitution might further expected that the species the same genus would occasionally exhibit reversions lost ancestral characters however never know the exact character the common ancestor group could not distinguish these two cases for instance did not know that the rock pigeon was not feather footed turn crowned could not have told whether these characters our domestic breeds were reversions only analogous variations but might have inferred that the blueness was case reversion from the number the markings which are correlated with the blue tint and which does not appear probable would all appear together from simple variation more especially might have inferred this from the blue colour and marks often appearing when distinct breeds diverse colours are crossed hence though under nature must generally left doubtful what cases are reversions anciently existing character and what are new but analogous variations yet ought theory sometimes find the varying offspring species assuming characters either from reversion from analogous variation which already occur some other members the same group and this undoubtedly the case nature considerable part the difficulty recognising variable species our systematic works due its varieties mocking were some the other species the same genus considerable catalogue also could given forms intermediate between two other forms which themselves must doubtfully ranked either varieties species and this shows unless all these forms considered independently created species that the one varying has assumed some the characters the other produce the intermediate form but the best evidence afforded parts organs important and uniform nature occasionally varying acquire some degree the character the same part organ allied species have collected long list such cases but here before lie under great disadvantage not being able give them can only repeat that such cases certainly occur and seem very remarkable will however give one curious and complex case not indeed affecting any important character but from occurring several species the same genus partly under domestication and partly under nature case apparently reversion the ass not rarely has very distinct transverse bars its legs like those the legs zebra has been asserted that these are plainest the foal and from inquiries which have made believe this true has also been asserted that the stripe each shoulder sometimes double the shoulder stripe certainly very variable length and outline white ass but not albino has been described without either spinal shoulder stripe and these stripes are sometimes very obscure actually quite lost dark coloured asses the koulan pallas said have been seen with double shoulder stripe the hemionus has shoulder stripe but traces stated blyth and others occasionally appear and have been informed colonel poole that the foals this species are generally striped the legs and faintly the shoulder the quagga though plainly barred like zebra over the body without bars the legs but gray has figured one specimen with very distinct zebra like bars the hocks with respect the horse have collected cases england the spinal stripe horses the most distinct breeds and all colours transverse bars the legs are not rare duns mouse duns and one instance chestnut faint shoulder stripe may sometimes seen duns and have seen trace bay horse son made careful examination and sketch for dun belgian cart horse with double stripe each shoulder and with leg stripes and man whom can implicitly trust has examined for small dun welch pony with three short parallel stripes each shoulder the north west part india the kattywar breed horses generally striped that hear from colonel poole who examined the breed for the indian government horse without stripes not considered purely bred the spine always striped the legs are generally barred and the shoulder stripe which sometimes double and sometimes treble common the side the face moreover sometimes striped the stripes are plainest the foal and sometimes quite disappear old horses colonel poole has seen both gray and bay kattywar horses striped when first foaled have also reason suspect from information given edwards that with the english race horse the spinal stripe much commoner the foal than the full grown animal without here entering further details may state that have collected cases leg and shoulder stripes horses very different breeds various countries from britain eastern china and from norway the north the malay archipelago the south all parts the world these stripes occur far oftenest duns and mouse duns the term dun large range colour included from one between brown and black close approach cream colour aware that colonel hamilton smith who has written this subject believes that the several breeds the horse have descended from several aboriginal species one which the dun was striped and that the above described appearances are all due ancient crosses with the dun stock but not all satisfied with this theory and should loth apply breeds distinct the heavy belgian cart horse welch ponies cobs the lanky kattywar race etc inhabiting the most distant parts the world now let turn the effects crossing the several species the horse genus rollin asserts that the common mule from the ass and horse particularly apt have bars its legs once saw mule with its legs much striped that any one first would have thought that must have been the product zebra and martin his excellent treatise the horse has given figure similar mule four coloured drawings which have seen hybrids between the ass and zebra the legs were much more plainly barred than the rest the body and one them there was double shoulder stripe lord moreton famous hybrid from chestnut mare and male quagga the hybrid and even the pure offspring subsequently produced from the mare black arabian sire were much more plainly barred across the legs than even the pure quagga lastly and this another most remarkable case hybrid has been figured gray and informs that knows second case from the ass and the hemionus and this hybrid though the ass seldom has stripes its legs and the hemionus has none and has not even shoulder stripe nevertheless had all four legs barred and had three short shoulder stripes like those the dun welch pony and even had some zebra like stripes the sides its face with respect this last fact was convinced that not even stripe colour appears from what would commonly called accident that was led solely from the occurrence the face stripes this hybrid from the ass and hemionus ask colonel poole whether such face stripes ever occur the eminently striped kattywar breed horses and was have seen answered the affirmative what now are say these several facts see several very distinct species the horse genus becoming simple variation striped the legs like zebra striped the shoulders like ass the horse see this tendency strong whenever dun tint appears tint which approaches that the general colouring the other species the genus the appearance the stripes not accompanied any change form any other new character see this tendency become striped most strongly displayed hybrids from between several the most distinct species now observe the case the several breeds pigeons they are descended from pigeon including two three sub species geographical races bluish colour with certain bars and other marks and when any breed assumes simple variation bluish tint these bars and other marks invariably reappear but without any other change form character when the oldest and truest breeds various colours are crossed see strong tendency for the blue tint and bars and marks reappear the mongrels have stated that the most probable hypothesis account for the reappearance very ancient characters that there tendency the young each successive generation produce the long lost character and that this tendency from unknown causes sometimes prevails and have just seen that several species the horse genus the stripes are either plainer appear more commonly the young than the old call the breeds pigeons some which have bred true for centuries species and how exactly parallel the case with that the species the horse genus for myself venture confidently look back thousands thousands generations and see animal striped like zebra but perhaps otherwise very differently constructed the common parent our domestic horse whether not descended from one more wild stocks the ass the hemionus quagga and zebra who believes that each equine species was independently created will presume assert that each species has been created with tendency vary both under nature and under domestication this particular manner often become striped like other species the genus and that each has been created with strong tendency when crossed with species inhabiting distant quarters the world produce hybrids resembling their stripes not their own parents but other species the genus admit this view seems reject real for unreal least for unknown cause makes the works god mere mockery and deception would almost soon believe with the old and ignorant cosmogonists that fossil shells had never lived but had been created stone mock the shells now living the sea shore summary our ignorance the laws variation profound not one case out hundred can pretend assign any reason why this that part differs more less from the same part the parents but whenever have the means instituting comparison the same laws appear have acted producing the lesser differences between varieties the same species and the greater differences between species the same genus the external conditions life climate and food etc seem have induced some slight modifications habit producing constitutional differences and use strengthening and disuse weakening and diminishing organs seem have been more potent their effects homologous parts tend vary the same way and homologous parts tend cohere modifications hard parts and external parts sometimes affect softer and internal parts when one part largely developed perhaps tends draw nourishment from the adjoining parts and every part the structure which can saved without detriment the individual will saved changes structure early age will generally affect parts subsequently developed and there are very many other correlations growth the nature which are utterly unable understand multiple parts are variable number and structure perhaps arising from such parts not having been closely specialised any particular function that their modifications have not been closely checked natural selection probably from this same cause that organic beings low the scale nature are more variable than those which have their whole organisation more specialised and are higher the scale rudimentary organs from being useless will disregarded natural selection and hence probably are variable specific characters that the characters which have come differ since the several species the same genus branched off from common parent are more variable than generic characters those which have long been inherited and have not differed within this same period these remarks have referred special parts organs being still variable because they have recently varied and thus come differ but have also seen the second chapter that the same principle applies the whole individual for district where many species any genus are found that where there has been much former variation and differentiation where the manufactory new specific forms has been actively work there average now find most varieties incipient species secondary sexual characters are highly variable and such characters differ much the species the same group variability the same parts the organisation has generally been taken advantage giving secondary sexual differences the sexes the same species and specific differences the several species the same genus any part organ developed extraordinary size extraordinary manner comparison with the same part organ the allied species must have gone through extraordinary amount modification since the genus arose and thus can understand why should often still variable much higher degree than other parts for variation long continued and slow process and natural selection will such cases not yet have had time overcome the tendency further variability and reversion less modified state but when species with any extraordinarily developed organ has become the parent many modified descendants which view must very slow process requiring long lapse time this case natural selection may readily have succeeded giving fixed character the organ however extraordinary manner may developed species inheriting nearly the same constitution from common parent and exposed similar influences will naturally tend present analogous variations and these same species may occasionally revert some the characters their ancient progenitors although new and important modifications may not arise from reversion and analogous variation such modifications will add the beautiful and harmonious diversity nature whatever the cause may each slight difference the offspring from their parents and cause for each must exist the steady accumulation through natural selection such differences when beneficial the individual that gives rise all the more important modifications structure which the innumerable beings the face this earth are enabled struggle with each other and the best adapted survive chapter difficulties theory difficulties the theory descent with modification transitions absence rarity transitional varieties transitions habits life diversified habits the same species species with habits widely different from those their allies organs extreme perfection means transition cases difficulty natura non facit saltum organs small importance organs not all cases absolutely perfect the law unity type and the conditions existence embraced the theory natural selection long before having arrived this part work crowd difficulties will have occurred the reader some them are grave that this day can never reflect them without being staggered but the best judgment the greater number are only apparent and those that are real are not think fatal theory these difficulties and objections may classed under the following heads firstly why species have descended from other species insensibly fine gradations not everywhere see innumerable transitional forms why not all nature confusion instead the species being see them well defined secondly possible that animal having for instance the structure and habits bat could have been formed the modification some animal with wholly different habits can believe that natural selection could produce the one hand organs trifling importance such the tail giraffe which serves fly flapper and the other hand organs such wonderful structure the eye which hardly yet fully understand the inimitable perfection thirdly can instincts acquired and modified through natural selection what shall say marvellous instinct that which leads the bee make cells which have practically anticipated the discoveries profound mathematicians fourthly how can account for species when crossed being sterile and producing sterile offspring whereas when varieties are crossed their fertility unimpaired the two first heads shall here discussed instinct and hybridism separate chapters the absence rarity transitional varieties natural selection acts solely the preservation profitable modifications each new form will tend fully stocked country take the place and finally exterminate its own less improved parent other less favoured forms with which comes into competition thus extinction and natural selection will have seen hand hand hence look each species descended from some other unknown form both the parent and all the transitional varieties will generally have been exterminated the very process formation and perfection the new form but this theory innumerable transitional forms must have existed why not find them embedded countless numbers the crust the earth will much more convenient discuss this question the chapter the imperfection the geological record and will here only state that believe the answer mainly lies the record being incomparably less perfect than generally supposed the imperfection the record being chiefly due organic beings not inhabiting profound depths the sea and their remains being embedded and preserved future age only masses sediment sufficiently thick and extensive withstand enormous amount future degradation and such fossiliferous masses can accumulated only where much sediment deposited the shallow bed the sea whilst slowly subsides these contingencies will concur only rarely and after enormously long intervals whilst the bed the sea stationary rising when very little sediment being deposited there will blanks our geological history the crust the earth vast museum but the natural collections have been made only intervals time immensely remote but may urged that when several closely allied species inhabit the same territory surely ought find the present time many transitional forms let take simple case travelling from north south over continent generally meet successive intervals with closely allied representative species evidently filling nearly the same place the natural economy the land these representative species often meet and interlock and the one becomes rarer and rarer the other becomes more and more frequent till the one replaces the other but compare these species where they intermingle they are generally absolutely distinct from each other every detail structure are specimens taken from the metropolis inhabited each theory these allied species have descended from common parent and during the process modification each has become adapted the conditions life its own region and has supplanted and exterminated its original parent and all the transitional varieties between its past and present states hence ought not expect the present time meet with numerous transitional varieties each region though they must have existed there and may embedded there fossil condition but the intermediate region having intermediate conditions life why not now find closely linking intermediate varieties this difficulty for long time quite confounded but think can large part explained the first place should extremely cautious inferring because area now continuous that has been continuous during long period geology would lead believe that almost every continent has been broken into islands even during the later tertiary periods and such islands distinct species might have been separately formed without the possibility intermediate varieties existing the intermediate zones changes the form the land and climate marine areas now continuous must often have existed within recent times far less continuous and uniform condition than present but will pass over this way escaping from the difficulty for believe that many perfectly defined species have been formed strictly continuous areas though not doubt that the formerly broken condition areas now continuous has played important part the formation new species more especially with freely crossing and wandering animals looking species they are now distributed over wide area generally find them tolerably numerous over large territory then becoming somewhat abruptly rarer and rarer the confines and finally disappearing hence the neutral territory between two representative species generally narrow comparison with the territory proper each see the same fact ascending mountains and sometimes quite remarkable how abruptly alph candolle has observed common alpine species disappears the same fact has been noticed forbes sounding the depths the sea with the dredge those who look climate and the physical conditions life the all important elements distribution these facts ought cause surprise climate and height depth graduate away insensibly but when bear mind that almost every species even its metropolis would increase immensely numbers were not for other competing species that nearly all either prey serve prey for others short that each organic being either directly indirectly related the most important manner other organic beings must see that the range the inhabitants any country means exclusively depends insensibly changing physical conditions but large part the presence other species which depends which destroyed with which comes into competition and these species are already defined objects however they may have become not blending one into another insensible gradations the range any one species depending does the range others will tend sharply defined moreover each species the confines its range where exists lessened numbers will during fluctuations the number its enemies its prey the seasons extremely liable utter extermination and thus its geographical range will come still more sharply defined right believing that allied representative species when inhabiting continuous area are generally distributed that each has wide range with comparatively narrow neutral territory between them which they become rather suddenly rarer and rarer then varieties not essentially differ from species the same rule will probably apply both and imagination adapt varying species very large area shall have adapt two varieties two large areas and third variety narrow intermediate zone the intermediate variety consequently will exist lesser numbers from inhabiting narrow and lesser area and practically far can make out this rule holds good with varieties state nature have met with striking instances the rule the case varieties intermediate between well marked varieties the genus balanus and would appear from information given watson asa gray and wollaston that generally when varieties intermediate between two other forms occur they are much rarer numerically than the forms which they connect now may trust these facts and inferences and therefore conclude that varieties linking two other varieties together have generally existed lesser numbers than the forms which they connect then think can understand why intermediate varieties should not endure for very long periods why general rule they should exterminated and disappear sooner than the forms which they originally linked together for any form existing lesser numbers would already remarked run greater chance being exterminated than one existing large numbers and this particular case the intermediate form would eminently liable the inroads closely allied forms existing both sides but far more important consideration believe that during the process further modification which two varieties are supposed theory converted and perfected into two distinct species the two which exist larger numbers from inhabiting larger areas will have great advantage over the intermediate variety which exists smaller numbers narrow and intermediate zone for forms existing larger numbers will always have better chance within any given period presenting further favourable variations for natural selection seize than will the rarer forms which exist lesser numbers hence the more common forms the race for life will tend beat and supplant the less common forms for these will more slowly modified and improved the same principle which believe accounts for the common species each country shown the second chapter presenting average greater number well marked varieties than the rarer species may illustrate what mean supposing three varieties sheep kept one adapted extensive mountainous region second comparatively narrow hilly tract and third wide plains the base and that the inhabitants are all trying with equal steadiness and skill improve their stocks selection the chances this case will strongly favour the great holders the mountains the plains improving their breeds more quickly than the small holders the intermediate narrow hilly tract and consequently the improved mountain plain breed will soon take the place the less improved hill breed and thus the two breeds which originally existed greater numbers will come into close contact with each other without the interposition the supplanted intermediate hill variety sum believe that species come tolerably well defined objects and not any one period present inextricable chaos varying and intermediate links firstly because new varieties are very slowly formed for variation very slow process and natural selection can nothing until favourable variations chance occur and until place the natural polity the country can better filled some modification some one more its inhabitants and such new places will depend slow changes climate the occasional immigration new inhabitants and probably still more important degree some the old inhabitants becoming slowly modified with the new forms thus produced and the old ones acting and reacting each other that any one region and any one time ought only see few species presenting slight modifications structure some degree permanent and this assuredly see secondly areas now continuous must often have existed within the recent period isolated portions which many forms more especially amongst the classes which unite for each birth and wander much may have separately been rendered sufficiently distinct rank representative species this case intermediate varieties between the several representative species and their common parent must formerly have existed each broken portion the land but these links will have been supplanted and exterminated during the process natural selection that they will longer exist living state thirdly when two more varieties have been formed different portions strictly continuous area intermediate varieties will probable first have been formed the intermediate zones but they will generally have had short duration for these intermediate varieties will from reasons already assigned namely from what know the actual distribution closely allied representative species and likewise acknowledged varieties exist the intermediate zones lesser numbers than the varieties which they tend connect from this cause alone the intermediate varieties will liable accidental extermination and during the process further modification through natural selection they will almost certainly beaten and supplanted the forms which they connect for these from existing greater numbers will the aggregate present more variation and thus further improved through natural selection and gain further advantages lastly looking not any one time but all time theory true numberless intermediate varieties linking most closely all the species the same group together must assuredly have existed but the very process natural selection constantly tends has been often remarked exterminate the parent forms and the intermediate links consequently evidence their former existence could found only amongst fossil remains which are preserved shall future chapter attempt show extremely imperfect and intermittent record the origin and transitions organic beings with peculiar habits and structure has been asked the opponents such views hold how for instance land carnivorous animal could have been converted into one with aquatic habits for how could the animal its transitional state have subsisted would easy show that within the same group carnivorous animals exist having every intermediate grade between truly aquatic and strictly terrestrial habits and each exists struggle for life clear that each well adapted its habits its place nature look the mustela vison north america which has webbed feet and which resembles otter its fur short legs and form tail during summer this animal dives for and preys fish but during the long winter leaves the frozen waters and preys like other polecats mice and land animals different case had been taken and had been asked how insectivorous quadruped could possibly have been converted into flying bat the question would have been far more difficult and could have given answer yet think such difficulties have very little weight here other occasions lie under heavy disadvantage for out the many striking cases which have collected can give only one two instances transitional habits and structures closely allied species the same genus and diversified habits either constant occasional the same species and seems that nothing less than long list such cases sufficient lessen the difficulty any particular case like that the bat look the family squirrels here have the finest gradation from animals with their tails only slightly flattened and from others sir richardson has remarked with the posterior part their bodies rather wide and with the skin their flanks rather full the called flying squirrels and flying squirrels have their limbs and even the base the tail united broad expanse skin which serves parachute and allows them glide through the air astonishing distance from tree tree cannot doubt that each structure use each kind squirrel its own country enabling escape birds beasts prey collect food more quickly there reason believe lessening the danger from occasional falls but does not follow from this fact that the structure each squirrel the best that possible conceive under all natural conditions let the climate and vegetation change let other competing rodents new beasts prey immigrate old ones become modified and all analogy would lead believe that some least the squirrels would decrease numbers become exterminated unless they also became modified and improved structure corresponding manner therefore can see difficulty more especially under changing conditions life the continued preservation individuals with fuller and fuller flank membranes each modification being useful each being propagated until the accumulated effects this process natural selection perfect called flying squirrel was produced now look the galeopithecus flying lemur which formerly was falsely ranked amongst bats has extremely wide flank membrane stretching from the corners the jaw the tail and including the limbs and the elongated fingers the flank membrane also furnished with extensor muscle although graduated links structure fitted for gliding through the air now connect the galeopithecus with the other lemuridæ yet can see difficulty supposing that such links formerly existed and that each had been formed the same steps the case the less perfectly gliding squirrels and that each grade structure had been useful its possessor nor can see any insuperable difficulty further believing possible that the membrane connected fingers and fore arm the galeopithecus might greatly lengthened natural selection and this far the organs flight are concerned would convert into bat bats which have the wing membrane extended from the top the shoulder the tail including the hind legs perhaps see traces apparatus originally constructed for gliding through the air rather than for flight about dozen genera birds had become extinct were unknown who would have ventured have surmised that birds might have existed which used their wings solely flappers like the logger headed duck micropterus eyton fins the water and front legs the land like the penguin sails like the ostrich and functionally for purpose like the apteryx yet the structure each these birds good for under the conditions life which exposed for each has live struggle but not necessarily the best possible under all possible conditions must not inferred from these remarks that any the grades wing structure here alluded which perhaps may all have resulted from disuse indicate the natural steps which birds have acquired their perfect power flight but they serve least show what diversified means transition are possible seeing that few members such water breathing classes the crustacea and mollusca are adapted live the land and seeing that have flying birds and mammals flying insects the most diversified types and formerly had flying reptiles conceivable that flying fish which now glide far through the air slightly rising and turning the aid their fluttering fins might have been modified into perfectly winged animals this had been effected who would have ever imagined that early transitional state they had been inhabitants the open ocean and had used their incipient organs flight exclusively far know escape being devoured other fish when see any structure highly perfected for any particular habit the wings bird for flight should bear mind that animals displaying early transitional grades the structure will seldom continue exist the present day for they will have been supplanted the very process perfection through natural selection furthermore may conclude that transitional grades between structures fitted for very different habits life will rarely have been developed early period great numbers and under many subordinate forms thus return our imaginary illustration the flying fish does not seem probable that fishes capable true flight would have been developed under many subordinate forms for taking prey many kinds many ways the land and the water until their organs flight had come high stage perfection have given them decided advantage over other animals the battle for life hence the chance discovering species with transitional grades structure fossil condition will always less from their having existed lesser numbers than the case species with fully developed structures will now give two three instances diversified and changed habits the individuals the same species when either case occurs would easy for natural selection fit the animal some modification its structure for its changed habits exclusively for one its several different habits but difficult tell and immaterial for whether habits generally change first and structure afterwards whether slight modifications structure lead changed habits both probably often change almost simultaneously cases changed habits will suffice merely allude that the many british insects which now feed exotic plants exclusively artificial substances diversified habits innumerable instances could given have often watched tyrant flycatcher saurophagus sulphuratus south america hovering over one spot and then proceeding another like kestrel and other times standing stationary the margin water and then dashing like kingfisher fish our own country the larger titmouse parus major may seen climbing branches almost like creeper often like shrike kills small birds blows the head and have many times seen and heard hammering the seeds the yew branch and thus breaking them like nuthatch north america the black bear was seen hearne swimming for hours with widely open mouth thus catching like whale insects the water even extreme case this the supply insects were constant and better adapted competitors did not already exist the country can see difficulty race bears being rendered natural selection more and more aquatic their structure and habits with larger and larger mouths till creature was produced monstrous whale sometimes see individuals species following habits widely different from those both their own species and the other species the same genus might expect theory that such individuals would occasionally have given rise new species having anomalous habits and with their structure either slightly considerably modified from that their proper type and such instances occur nature can more striking instance adaptation given than that woodpecker for climbing trees and for seizing insects the chinks the bark yet north america there are woodpeckers which feed largely fruit and others with elongated wings which chase insects the wing and the plains plata where not tree grows there woodpecker which every essential part its organisation even its colouring the harsh tone its voice and undulatory flight told plainly its close blood relationship our common species yet woodpecker which never climbs tree petrels are the most aërial and oceanic birds yet the quiet sounds tierra del fuego the puffinuria berardi its general habits its astonishing power diving its manner swimming and flying when unwillingly takes flight would mistaken any one for auk grebe nevertheless essentially petrel but with many parts its organisation profoundly modified the other hand the acutest observer examining the dead body the water ouzel would never have suspected its sub aquatic habits yet this anomalous member the strictly terrestrial thrush family wholly subsists diving grasping the stones with its feet and using its wings under water who believes that each being has been created now see must occasionally have felt surprise when has met with animal having habits and structure not all agreement what can plainer than that the webbed feet ducks and geese are formed for swimming yet there are upland geese with webbed feet which rarely never near the water and one except audubon has seen the frigate bird which has all its four toes webbed alight the surface the sea the other hand grebes and coots are eminently aquatic although their toes are only bordered membrane what seems plainer than that the long toes grallatores are formed for walking over swamps and floating plants yet the water hen nearly aquatic the coot and the landrail nearly terrestrial the quail partridge such cases and many others could given habits have changed without corresponding change structure the webbed feet the upland goose may said have become rudimentary function though not structure the frigate bird the deeply scooped membrane between the toes shows that structure has begun change who believes separate and innumerable acts creation will say that these cases has pleased the creator cause being one type take the place one another type but this seems only restating the fact dignified language who believes the struggle for existence and the principle natural selection will acknowledge that every organic being constantly endeavouring increase numbers and that any one being vary ever little either habits structure and thus gain advantage over some other inhabitant the country will seize the place that inhabitant however different may from its own place hence will cause him surprise that there should geese and frigate birds with webbed feet either living the dry land most rarely alighting the water that there should long toed corncrakes living meadows instead swamps that there should woodpeckers where not tree grows that there should diving thrushes and petrels with the habits auks organs extreme perfection and complication suppose that the eye with all its inimitable contrivances for adjusting the focus different distances for admitting different amounts light and for the correction spherical and chromatic aberration could have been formed natural selection seems freely confess absurd the highest possible degree yet reason tells that numerous gradations from perfect and complex eye one very imperfect and simple each grade being useful its possessor can shown exist further the eye does vary ever slightly and the variations inherited which certainly the case and any variation modification the organ ever useful animal under changing conditions life then the difficulty believing that perfect and complex eye could formed natural selection though insuperable our imagination can hardly considered real how nerve comes sensitive light hardly concerns more than how life itself first originated but may remark that several facts make suspect that any sensitive nerve may rendered sensitive light and likewise those coarser vibrations the air which produce sound looking for the gradations which organ any species has been perfected ought look exclusively its lineal ancestors but this scarcely ever possible and are forced each case look species the same group that the collateral descendants from the same original parent form order see what gradations are possible and for the chance some gradations having been transmitted from the earlier stages descent unaltered little altered condition amongst existing vertebrata find but small amount gradation the structure the eye and from fossil species can learn nothing this head this great class should probably have descend far beneath the lowest known fossiliferous stratum discover the earlier stages which the eye has been perfected the articulata can commence series with optic nerve merely coated with pigment and without any other mechanism and from this low stage numerous gradations structure branching off two fundamentally different lines can shown exist until reach moderately high stage perfection certain crustaceans for instance there double cornea the inner one divided into facets within each which there lens shaped swelling other crustaceans the transparent cones which are coated pigment and which properly act only excluding lateral pencils light are convex their upper ends and must act convergence and their lower ends there seems imperfect vitreous substance with these facts here far too briefly and imperfectly given which show that there much graduated diversity the eyes living crustaceans and bearing mind how small the number living animals proportion those which have become extinct can see very great difficulty not more than the case many other structures believing that natural selection has converted the simple apparatus optic nerve merely coated with pigment and invested transparent membrane into optical instrument perfect possessed any member the great articulate class who will thus far find finishing this treatise that large bodies facts otherwise inexplicable can explained the theory descent ought not hesitate further and admit that structure even perfect the eye eagle might formed natural selection although this case does not know any the transitional grades his reason ought conquer his imagination though have felt the difficulty far too keenly surprised any degree hesitation extending the principle natural selection such startling lengths scarcely possible avoid comparing the eye telescope know that this instrument has been perfected the long continued efforts the highest human intellects and naturally infer that the eye has been formed somewhat analogous process but may not this inference presumptuous have any right assume that the creator works intellectual powers like those man must compare the eye optical instrument ought imagination take thick layer transparent tissue with nerve sensitive light beneath and then suppose every part this layer continually changing slowly density separate into layers different densities and thicknesses placed different distances from each other and with the surfaces each layer slowly changing form further must suppose that there power always intently watching each slight accidental alteration the transparent layers and carefully selecting each alteration which under varied circumstances may any way any degree tend produce distincter image must suppose each new state the instrument multiplied the million and each preserved till better produced and then the old ones destroyed living bodies variation will cause the slight alterations generation will multiply them almost infinitely and natural selection will pick out with unerring skill each improvement let this process for millions millions years and during each year millions individuals many kinds and may not believe that living optical instrument might thus formed superior one glass the works the creator are those man could demonstrated that any complex organ existed which could not possibly have been formed numerous successive slight modifications theory would absolutely break down but can find out such case doubt many organs exist which not know the transitional grades more especially look much isolated species round which according theory there has been much extinction again look organ common all the members large class for this latter case the organ must have been first formed extremely remote period since which all the many members the class have been developed and order discover the early transitional grades through which the organ has passed should have look very ancient ancestral forms long since become extinct should extremely cautious concluding that organ could not have been formed transitional gradations some kind numerous cases could given amongst the lower animals the same organ performing the same time wholly distinct functions thus the alimentary canal respires digests and excretes the larva the dragon fly and the fish cobites the hydra the animal may turned inside out and the exterior surface will then digest and the stomach respire such cases natural selection might easily specialise any advantage were thus gained part organ which had performed two functions for one function alone and thus wholly change its nature insensible steps two distinct organs sometimes perform simultaneously the same function the same individual give one instance there are fish with gills branchiæ that breathe the air dissolved the water the same time that they breathe free air their swimbladders this latter organ having ductus pneumaticus for its supply and being divided highly vascular partitions these cases one the two organs might with ease modified and perfected perform all the work itself being aided during the process modification the other organ and then this other organ might modified for some other and quite distinct purpose quite obliterated the illustration the swimbladder fishes good one because shows clearly the highly important fact that organ originally constructed for one purpose namely flotation may converted into one for wholly different purpose namely respiration the swimbladder has also been worked accessory the auditory organs certain fish for not know which view now generally held part the auditory apparatus has been worked complement the swimbladder all physiologists admit that the swimbladder homologous ideally similar position and structure with the lungs the higher vertebrate animals hence there seems great difficulty believing that natural selection has actually converted swimbladder into lung organ used exclusively for respiration can indeed hardly doubt that all vertebrate animals having true lungs have descended ordinary generation from ancient prototype which know nothing furnished with floating apparatus swimbladder can thus infer from professor owen interesting description these parts understand the strange fact that every particle food and drink which swallow has pass over the orifice the trachea with some risk falling into the lungs notwithstanding the beautiful contrivance which the glottis closed the higher vertebrata the branchiæ have wholly disappeared the slits the sides the neck and the loop like course the arteries still marking the embryo their former position but conceivable that the now utterly lost branchiæ might have been gradually worked natural selection for some quite distinct purpose the same manner the view entertained some naturalists that the branchiæ and dorsal scales annelids are homologous with the wings and wing covers insects probable that organs which very ancient period served for respiration have been actually converted into organs flight considering transitions organs important bear mind the probability conversion from one function another that will give one more instance pedunculated cirripedes have two minute folds skin called the ovigerous frena which serve through the means sticky secretion retain the eggs until they are hatched within the sack these cirripedes have branchiæ the whole surface the body and sack including the small frena serving for respiration the balanidæ sessile cirripedes the other hand have ovigerous frena the eggs lying loose the bottom the sack the well enclosed shell but they have large folded branchiæ now think one will dispute that the ovigerous frena the one family are strictly homologous with the branchiæ the other family indeed they graduate into each other therefore not doubt that little folds skin which originally served ovigerous frena but which likewise very slightly aided the act respiration have been gradually converted natural selection into branchiæ simply through increase their size and the obliteration their adhesive glands all pedunculated cirripedes had become extinct and they have already suffered far more extinction than have sessile cirripedes who would ever have imagined that the branchiæ this latter family had originally existed organs for preventing the ova from being washed out the sack although must extremely cautious concluding that any organ could not possibly have been produced successive transitional gradations yet undoubtedly grave cases difficulty occur some which will discussed future work one the gravest that neuter insects which are often very differently constructed from either the males fertile females but this case will treated the next chapter the electric organs fishes offer another case special difficulty impossible conceive what steps these wondrous organs have been produced but owen and others have remarked their intimate structure closely resembles that common muscle and has lately been shown that rays have organ closely analogous the electric apparatus and yet not matteuchi asserts discharge any electricity must own that are far too ignorant argue that transition any kind possible the electric organs offer another and even more serious difficulty for they occur only about dozen fishes which several are widely remote their affinities generally when the same organ appears several members the same class especially members having very different habits life may attribute its presence inheritance from common ancestor and its absence some the members its loss through disuse natural selection but the electric organs had been inherited from one ancient progenitor thus provided might have expected that all electric fishes would have been specially related each other nor does geology all lead the belief that formerly most fishes had electric organs which most their modified descendants have lost the presence luminous organs few insects belonging different families and orders offers parallel case difficulty other cases could given for instance plants the very curious contrivance mass pollen grains borne foot stalk with sticky gland the end the same orchis and asclepias genera almost remote possible amongst flowering plants all these cases two very distinct species furnished with apparently the same anomalous organ should observed that although the general appearance and function the organ may the same yet some fundamental difference can generally detected inclined believe that nearly the same way two men have sometimes independently hit the very same invention natural selection working for the good each being and taking advantage analogous variations has sometimes modified very nearly the same manner two parts two organic beings which owe but little their structure common inheritance from the same ancestor although many cases most difficult conjecture what transitions organ could have arrived its present state yet considering that the proportion living and known forms the extinct and unknown very small have been astonished how rarely organ can named towards which transitional grade known lead the truth this remark indeed shown that old canon natural history natura non facit saltum meet with this admission the writings almost every experienced naturalist milne edwards has well expressed nature prodigal variety but niggard innovation why the theory creation should this why should all the parts and organs many independent beings each supposed have been separately created for its proper place nature invariably linked together graduated steps why should not nature have taken leap from structure structure the theory natural selection can clearly understand why she should not for natural selection can act only taking advantage slight successive variations she can never take leap but must advance the shortest and slowest steps organs little apparent importance natural selection acts life and death the preservation individuals with any favourable variation and the destruction those with any unfavourable deviation structure have sometimes felt much difficulty understanding the origin simple parts which the importance does not seem sufficient cause the preservation successively varying individuals have sometimes felt much difficulty though very different kind this head the case organ perfect and complex the eye the first place are much too ignorant regard the whole economy any one organic being say what slight modifications would importance not former chapter have given instances most trifling characters such the down fruit and the colour the flesh which from determining the attacks insects from being correlated with constitutional differences might assuredly acted natural selection the tail the giraffe looks like artificially constructed fly flapper and seems first incredible that this could have been adapted for its present purpose successive slight modifications each better and better for trifling object driving away flies yet should pause before being too positive even this case for know that the distribution and existence cattle and other animals south america absolutely depends their power resisting the attacks insects that individuals which could any means defend themselves from these small enemies would able range into new pastures and thus gain great advantage not that the larger quadrupeds are actually destroyed except some rare cases the flies but they are incessantly harassed and their strength reduced that they are more subject disease not well enabled coming dearth search for food escape from beasts prey organs now trifling importance have probably some cases been high importance early progenitor and after having been slowly perfected former period have been transmitted nearly the same state although now become very slight use and any actually injurious deviations their structure will always have been checked natural selection seeing how important organ locomotion the tail most aquatic animals its general presence and use for many purposes many land animals which their lungs modified swim bladders betray their aquatic origin may perhaps thus accounted for well developed tail having been formed aquatic animal might subsequently come worked for all sorts purposes fly flapper organ prehension aid turning with the dog though the aid must slight for the hare with hardly any tail can double quickly enough the second place may sometimes attribute importance characters which are really very little importance and which have originated from quite secondary causes independently natural selection should remember that climate food etc probably have some little direct influence the organisation that characters reappear from the law reversion that correlation growth will have had most important influence modifying various structures and finally that sexual selection will often have largely modified the external characters animals having will give one male advantage fighting with another charming the females moreover when modification structure has primarily arisen from the above other unknown causes may first have been advantage the species but may subsequently have been taken advantage the descendants the species under new conditions life and with newly acquired habits give few instances illustrate these latter remarks green woodpeckers alone had existed and did not know that there were many black and pied kinds dare say that should have thought that the green colour was beautiful adaptation hide this tree frequenting bird from its enemies and consequently that was character importance and might have been acquired through natural selection have doubt that the colour due some quite distinct cause probably sexual selection trailing bamboo the malay archipelago climbs the loftiest trees the aid exquisitely constructed hooks clustered around the ends the branches and this contrivance doubt the highest service the plant but see nearly similar hooks many trees which are not climbers the hooks the bamboo may have arisen from unknown laws growth and have been subsequently taken advantage the plant undergoing further modification and becoming climber the naked skin the head vulture generally looked direct adaptation for wallowing putridity and may may possibly due the direct action putrid matter but should very cautious drawing any such inference when see that the skin the head the clean feeding male turkey likewise naked the sutures the skulls young mammals have been advanced beautiful adaptation for aiding parturition and doubt they facilitate may indispensable for this act but sutures occur the skulls young birds and reptiles which have only escape from broken egg may infer that this structure has arisen from the laws growth and has been taken advantage the parturition the higher animals are profoundly ignorant the causes producing slight and unimportant variations and are immediately made conscious this reflecting the differences the breeds our domesticated animals different countries more especially the less civilized countries where there has been but little artificial selection careful observers are convinced that damp climate affects the growth the hair and that with the hair the horns are correlated mountain breeds always differ from lowland breeds and mountainous country would probably affect the hind limbs from exercising them more and possibly even the form the pelvis and then the law homologous variation the front limbs and even the head would probably affected the shape also the pelvis might affect pressure the shape the head the young the womb the laborious breathing necessary high regions would have some reason believe increase the size the chest and again correlation would come into play animals kept savages different countries often have struggle for their own subsistence and would exposed certain extent natural selection and individuals with slightly different constitutions would succeed best under different climates and there reason believe that constitution and colour are correlated good observer also states that cattle susceptibility the attacks flies correlated with colour the liability poisoned certain plants that colour would thus subjected the action natural selection but are far too ignorant speculate the relative importance the several known and unknown laws variation and have here alluded them only show that are unable account for the characteristic differences our domestic breeds which nevertheless generally admit have arisen through ordinary generation ought not lay too much stress our ignorance the precise cause the slight analogous differences between species might have adduced for this same purpose the differences between the races man which are strongly marked may add that some little light can apparently thrown the origin these differences chiefly through sexual selection particular kind but without here entering copious details reasoning would appear frivolous the foregoing remarks lead say few words the protest lately made some naturalists against the utilitarian doctrine that every detail structure has been produced for the good its possessor they believe that very many structures have been created for beauty the eyes man for mere variety this doctrine true would absolutely fatal theory yet fully admit that many structures are direct use their possessors physical conditions probably have had some little effect structure quite independently any good thus gained correlation growth has doubt played most important part and useful modification one part will often have entailed other parts diversified changes direct use again characters which formerly were useful which formerly had arisen from correlation growth from other unknown cause may reappear from the law reversion though now direct use the effects sexual selection when displayed beauty charm the females can called useful only rather forced sense but far the most important consideration that the chief part the organisation every being simply due inheritance and consequently though each being assuredly well fitted for its place nature many structures now have direct relation the habits life each species thus can hardly believe that the webbed feet the upland goose the frigate bird are special use these birds cannot believe that the same bones the arm the monkey the fore leg the horse the wing the bat and the flipper the seal are special use these animals may safely attribute these structures inheritance but the progenitor the upland goose and the frigate bird webbed feet doubt were useful they now are the most aquatic existing birds may believe that the progenitor the seal had not flipper but foot with five toes fitted for walking grasping and may further venture believe that the several bones the limbs the monkey horse and bat which have been inherited from common progenitor were formerly more special use that progenitor its progenitors than they now are these animals having such widely diversified habits therefore may infer that these several bones might have been acquired through natural selection subjected formerly now the several laws inheritance reversion correlation growth etc hence every detail structure every living creature making some little allowance for the direct action physical conditions may viewed either having been special use some ancestral form being now special use the descendants this form either directly indirectly through the complex laws growth natural selection cannot possibly produce any modification any one species exclusively for the good another species though throughout nature one species incessantly takes advantage and profits the structure another but natural selection can and does often produce structures for the direct injury other species see the fang the adder and the ovipositor the ichneumon which its eggs are deposited the living bodies other insects could proved that any part the structure any one species had been formed for the exclusive good another species would annihilate theory for such could not have been produced through natural selection although many statements may found works natural history this effect cannot find even one which seems any weight admitted that the rattlesnake has poison fang for its own defence and for the destruction its prey but some authors suppose that the same time this snake furnished with rattle for its own injury namely warn its prey escape would almost soon believe that the cat curls the end its tail when preparing spring order warn the doomed mouse but have not space here enter this and other such cases natural selection will never produce being anything injurious itself for natural selection acts solely and for the good each organ will formed paley has remarked for the purpose causing pain for doing injury its possessor fair balance struck between the good and evil caused each part each will found the whole advantageous after the lapse time under changing conditions life any part comes injurious will modified not the being will become extinct myriads have become extinct natural selection tends only make each organic being perfect slightly more perfect than the other inhabitants the same country with which has struggle for existence and see that this the degree perfection attained under nature the endemic productions new zealand for instance are perfect one compared with another but they are now rapidly yielding before the advancing legions plants and animals introduced from europe natural selection will not produce absolute perfection nor always meet far can judge with this high standard under nature the correction for the aberration light said high authority not perfect even that most perfect organ the eye our reason leads admire with enthusiasm multitude inimitable contrivances nature this same reason tells though may easily err both sides that some other contrivances are less perfect can consider the sting the wasp the bee perfect which when used against many attacking animals cannot withdrawn owing the backward serratures and inevitably causes the death the insect tearing out its viscera look the sting the bee having originally existed remote progenitor boring and serrated instrument like that many members the same great order and which has been modified but not perfected for its present purpose with the poison originally adapted cause galls subsequently intensified can perhaps understand how that the use the sting should often cause the insect own death for the whole the power stinging useful the community will fulfil all the requirements natural selection though may cause the death some few members admire the truly wonderful power scent which the males many insects find their females can admire the production for this single purpose thousands drones which are utterly useless the community for any other end and which are ultimately slaughtered their industrious and sterile sisters may difficult but ought admire the savage instinctive hatred the queen bee which urges her instantly destroy the young queens her daughters soon born perish herself the combat for undoubtedly this for the good the community and maternal love maternal hatred though the latter fortunately most rare all the same the inexorable principle natural selection admire the several ingenious contrivances which the flowers the orchis and many other plants are fertilised through insect agency can consider equally perfect the elaboration our fir trees dense clouds pollen order that few granules may wafted chance breeze the ovules summary chapter have this chapter discussed some the difficulties and objections which may urged against theory many them are very grave but think that the discussion light has been thrown several facts which the theory independent acts creation are utterly obscure have seen that species any one period are not indefinitely variable and are not linked together multitude intermediate gradations partly because the process natural selection will always very slow and will act any one time only very few forms and partly because the very process natural selection almost implies the continual supplanting and extinction preceding and intermediate gradations closely allied species now living continuous area must often have been formed when the area was not continuous and when the conditions life did not insensibly graduate away from one part another when two varieties are formed two districts continuous area intermediate variety will often formed fitted for intermediate zone but from reasons assigned the intermediate variety will usually exist lesser numbers than the two forms which connects consequently the two latter during the course further modification from existing greater numbers will have great advantage over the less numerous intermediate variety and will thus generally succeed supplanting and exterminating have seen this chapter how cautious should concluding that the most different habits life could not graduate into each other that bat for instance could not have been formed natural selection from animal which first could only glide through the air have seen that species may under new conditions life change its habits have diversified habits with some habits very unlike those its nearest congeners hence can understand bearing mind that each organic being trying live wherever can live how has arisen that there are upland geese with webbed feet ground woodpeckers diving thrushes and petrels with the habits auks although the belief that organ perfect the eye could have been formed natural selection more than enough stagger any one yet the case any organ know long series gradations complexity each good for its possessor then under changing conditions life there logical impossibility the acquirement any conceivable degree perfection through natural selection the cases which know intermediate transitional states should very cautious concluding that none could have existed for the homologies many organs and their intermediate states show that wonderful metamorphoses function are least possible for instance swim bladder has apparently been converted into air breathing lung the same organ having performed simultaneously very different functions and then having been specialised for one function and two very distinct organs having performed the same time the same function the one having been perfected whilst aided the other must often have largely facilitated transitions are far too ignorant almost every case enabled assert that any part organ unimportant for the welfare species that modifications its structure could not have been slowly accumulated means natural selection but may confidently believe that many modifications wholly due the laws growth and first way advantageous species have been subsequently taken advantage the still further modified descendants this species may also believe that part formerly high importance has often been retained the tail aquatic animal its terrestrial descendants though has become such small importance that could not its present state have been acquired natural selection power which acts solely the preservation profitable variations the struggle for life natural selection will produce nothing one species for the exclusive good injury another though may well produce parts organs and excretions highly useful even indispensable highly injurious another species but all cases the same time useful the owner natural selection each well stocked country must act chiefly through the competition the inhabitants one with another and consequently will produce perfection strength the battle for life only according the standard that country hence the inhabitants one country generally the smaller one will often yield see they yield the inhabitants another and generally larger country for the larger country there will have existed more individuals and more diversified forms and the competition will have been severer and thus the standard perfection will have been rendered higher natural selection will not necessarily produce absolute perfection nor far can judge our limited faculties can absolute perfection everywhere found the theory natural selection can clearly understand the full meaning that old canon natural history natura non facit saltum this canon look only the present inhabitants the world not strictly correct but include all those past times must theory strictly true generally acknowledged that all organic beings have been formed two great laws unity type and the conditions existence unity type meant that fundamental agreement structure which see organic beings the same class and which quite independent their habits life theory unity type explained unity descent the expression conditions existence often insisted the illustrious cuvier fully embraced the principle natural selection for natural selection acts either now adapting the varying parts each being its organic and inorganic conditions life having adapted them during long past periods time the adaptations being aided some cases use and disuse being slightly affected the direct action the external conditions life and being all cases subjected the several laws growth hence fact the law the conditions existence the higher law includes through the inheritance former adaptations that unity type chapter vii instinct instincts comparable with habits but different their origin instincts graduated aphides and ants instincts variable domestic instincts their origin natural instincts the cuckoo ostrich and parasitic bees slave making ants hive bee its cell making instinct difficulties the theory the natural selection instincts neuter sterile insects summary the subject instinct might have been worked into the previous chapters but have thought that would more convenient treat the subject separately especially wonderful instinct that the hive bee making its cells will probably have occurred many readers difficulty sufficient overthrow whole theory must premise that have nothing with the origin the primary mental powers any more than have with that life itself are concerned only with the diversities instinct and the other mental qualities animals within the same class will not attempt any definition instinct would easy show that several distinct mental actions are commonly embraced this term but every one understands what meant when said that instinct impels the cuckoo migrate and lay her eggs other birds nests action which ourselves should require experience enable perform when performed animal more especially very young one without any experience and when performed many individuals the same way without their knowing for what purpose performed usually said instinctive but could show that none these characters instinct are universal little dose pierre huber expresses judgment reason often comes into play even animals very low the scale nature frederick cuvier and several the older metaphysicians have compared instinct with habit this comparison gives think remarkably accurate notion the frame mind under which instinctive action performed but not its origin how unconsciously many habitual actions are performed indeed not rarely direct opposition our conscious will yet they may modified the will reason habits easily become associated with other habits and with certain periods time and states the body when once acquired they often remain constant throughout life several other points resemblance between instincts and habits could pointed out repeating well known song instincts one action follows another sort rhythm person interrupted song repeating anything rote generally forced back recover the habitual train thought huber found was with caterpillar which makes very complicated hammock for took caterpillar which had completed its hammock say the sixth stage construction and put into hammock completed only the third stage the caterpillar simply performed the fourth fifth and sixth stages construction however caterpillar were taken out hammock made for instance the third stage and were put into one finished the sixth stage that much its work was already done for far from feeling the benefit this was much embarrassed and order complete its hammock seemed forced start from the third stage where had left off and thus tried complete the already finished work suppose any habitual action become inherited and think can shown that this does sometimes happen then the resemblance between what originally was habit and instinct becomes close not distinguished mozart instead playing the pianoforte three years old with wonderfully little practice had played tune with practice all might truly said have done instinctively but would the most serious error suppose that the greater number instincts have been acquired habit one generation and then transmitted inheritance succeeding generations can clearly shown that the most wonderful instincts with which are acquainted namely those the hive bee and many ants could not possibly have been thus acquired will universally admitted that instincts are important corporeal structure for the welfare each species under its present conditions life under changed conditions life least possible that slight modifications instinct might profitable species and can shown that instincts vary ever little then can see difficulty natural selection preserving and continually accumulating variations instinct any extent that may profitable thus believe that all the most complex and wonderful instincts have originated modifications corporeal structure arise from and are increased use habit and are diminished lost disuse not doubt has been with instincts but believe that the effects habit are quite subordinate importance the effects the natural selection what may called accidental variations instincts that variations produced the same unknown causes which produce slight deviations bodily structure complex instinct can possibly produced through natural selection except the slow and gradual accumulation numerous slight yet profitable variations hence the case corporeal structures ought find nature not the actual transitional gradations which each complex instinct has been acquired for these could found only the lineal ancestors each species but ought find the collateral lines descent some evidence such gradations ought least able show that gradations some kind are possible and this certainly can have been surprised find making allowance for the instincts animals having been but little observed except europe and north america and for instinct being known amongst extinct species how very generally gradations leading the most complex instincts can discovered the canon natura non facit saltum applies with almost equal force instincts bodily organs changes instinct may sometimes facilitated the same species having different instincts different periods life different seasons the year when placed under different circumstances etc which case either one the other instinct might preserved natural selection and such instances diversity instinct the same species can shown occur nature again the case corporeal structure and conformably with theory the instinct each species good for itself but has never far can judge been produced for the exclusive good others one the strongest instances animal apparently performing action for the sole good another with which acquainted that aphides voluntarily yielding their sweet excretion ants that they voluntarily the following facts show removed all the ants from group about dozen aphides dock plant and prevented their attendance during several hours after this interval felt sure that the aphides would want excrete watched them for some time through lens but not one excreted then tickled and stroked them with hair the same manner well could the ants with their antennæ but not one excreted afterwards allowed ant visit them and immediately seemed its eager way running about well aware what rich flock had discovered then began play with its antennæ the abdomen first one aphis and then another and each aphis soon felt the antennæ immediately lifted its abdomen and excreted limpid drop sweet juice which was eagerly devoured the ant even the quite young aphides behaved this manner showing that the action was instinctive and not the result experience but the excretion extremely viscid probably convenience the aphides have removed and therefore probably the aphides not instinctively excrete for the sole good the ants although not believe that any animal the world performs action for the exclusive good another distinct species yet each species tries take advantage the instincts others each takes advantage the weaker bodily structure others again some few cases certain instincts cannot considered absolutely perfect but details this and other such points are not indispensable they may here passed over some degree variation instincts under state nature and the inheritance such variations are indispensable for the action natural selection many instances possible ought have been here given but want space prevents can only assert that instincts certainly vary for instance the migratory instinct both extent and direction and its total loss with the nests birds which vary partly dependence the situations chosen and the nature and temperature the country inhabited but often from causes wholly unknown audubon has given several remarkable cases differences nests the same species the northern and southern united states fear any particular enemy certainly instinctive quality may seen nestling birds though strengthened experience and the sight fear the same enemy other animals but fear man slowly acquired have elsewhere shown various animals inhabiting desert islands and may see instance this even england the greater wildness all our large birds than our small birds for the large birds have been most persecuted man may safely attribute the greater wildness our large birds this cause for uninhabited islands large birds are not more fearful than small and the magpie wary england tame norway the hooded crow egypt that the general disposition individuals the same species born state nature extremely diversified can shown multitude facts several cases also could given occasional and strange habits certain species which might advantageous the species give rise through natural selection quite new instincts but well aware that these general statements without facts given detail can produce but feeble effect the reader mind can only repeat assurance that not speak without good evidence the possibility even probability inherited variations instinct state nature will strengthened briefly considering few cases under domestication shall thus also enabled see the respective parts which habit and the selection called accidental variations have played modifying the mental qualities our domestic animals number curious and authentic instances could given the inheritance all shades disposition and tastes and likewise the oddest tricks associated with certain frames mind periods time but let look the familiar case the several breeds dogs cannot doubted that young pointers have myself seen striking instance will sometimes point and even back other dogs the very first time that they are taken out retrieving certainly some degree inherited retrievers and tendency run round instead flock sheep shepherd dogs cannot see that these actions performed without experience the young and nearly the same manner each individual performed with eager delight each breed and without the end being known for the young pointer can more know that points aid his master than the white butterfly knows why she lays her eggs the leaf the cabbage cannot see that these actions differ essentially from true instincts were see one kind wolf when young and without any training soon scented its prey stand motionless like statue and then slowly crawl forward with peculiar gait and another kind wolf rushing round instead herd deer and driving them distant point should assuredly call these actions instinctive domestic instincts they may called are certainly far less fixed invariable than natural instincts but they have been acted far less rigorous selection and have been transmitted for incomparably shorter period under less fixed conditions life how strongly these domestic instincts habits and dispositions are inherited and how curiously they become mingled well shown when different breeds dogs are crossed thus known that cross with bull dog has affected for many generations the courage and obstinacy greyhounds and cross with greyhound has given whole family shepherd dogs tendency hunt hares these domestic instincts when thus tested crossing resemble natural instincts which like manner become curiously blended together and for long period exhibit traces the instincts either parent for example roy describes dog whose great grandfather was wolf and this dog showed trace its wild parentage only one way not coming straight line his master when called domestic instincts are sometimes spoken actions which have become inherited solely from long continued and compulsory habit but this think not true one would ever have thought teaching probably could have taught the tumbler pigeon tumble action which have witnessed performed young birds that have never seen pigeon tumble may believe that some one pigeon showed slight tendency this strange habit and that the long continued selection the best individuals successive generations made tumblers what they now are and near glasgow there are house tumblers hear from brent which cannot fly eighteen inches high without going head over heels may doubted whether any one would have thought training dog point had not some one dog naturally shown tendency this line and this known occasionally happen once saw pure terrier when the first tendency was once displayed methodical selection and the inherited effects compulsory training each successive generation would soon complete the work and unconscious selection still work each man tries procure without intending improve the breed dogs which will stand and hunt best the other hand habit alone some cases has sufficed animal more difficult tame than the young the wild rabbit scarcely any animal tamer than the young the tame rabbit but not suppose that domestic rabbits have ever been selected for tameness and presume that must attribute the whole the inherited change from extreme wildness extreme tameness simply habit and long continued close confinement natural instincts are lost under domestication remarkable instance this seen those breeds fowls which very rarely never become broody that never wish sit their eggs familiarity alone prevents our seeing how universally and largely the minds our domestic animals have been modified domestication scarcely possible doubt that the love man has become instinctive the dog all wolves foxes jackals and species the cat genus when kept tame are most eager attack poultry sheep and pigs and this tendency has been found incurable dogs which have been brought home puppies from countries such tierra del fuego and australia where the savages not keep these domestic animals how rarely the other hand our civilised dogs even when quite young require taught not attack poultry sheep and pigs doubt they occasionally make attack and are then beaten and not cured they are destroyed that habit with some degree selection has probably concurred civilising inheritance our dogs the other hand young chickens have lost wholly habit that fear the dog and cat which doubt was originally instinctive them the same way plainly instinctive young pheasants though reared under hen not that chickens have lost all fear but fear only dogs and cats for the hen gives the danger chuckle they will run more especially young turkeys from under her and conceal themselves the surrounding grass thickets and this evidently done for the instinctive purpose allowing see wild ground birds their mother fly away but this instinct retained our chickens has become useless under domestication for the mother hen has almost lost disuse the power flight hence may conclude that domestic instincts have been acquired and natural instincts have been lost partly habit and partly man selecting and accumulating during successive generations peculiar mental habits and actions which first appeared from what must our ignorance call accident some cases compulsory habit alone has sufficed produce such inherited mental changes other cases compulsory habit has done nothing and all has been the result selection pursued both methodically and unconsciously but most cases probably habit and selection have acted together shall perhaps best understand how instincts state nature have become modified selection considering few cases will select only three out the several which shall have discuss future work namely the instinct which leads the cuckoo lay her eggs other birds nests the slave making instinct certain ants and the comb making power the hive bee these two latter instincts have generally and most justly been ranked naturalists the most wonderful all known instincts now commonly admitted that the more immediate and final cause the cuckoo instinct that she lays her eggs not daily but intervals two three days that she were make her own nest and sit her own eggs those first laid would have left for some time unincubated there would eggs and young birds different ages the same nest this were the case the process laying and hatching might inconveniently long more especially she has migrate very early period and the first hatched young would probably have fed the male alone but the american cuckoo this predicament for she makes her own nest and has eggs and young successively hatched all the same time has been asserted that the american cuckoo occasionally lays her eggs other birds nests but hear the high authority brewer that this mistake nevertheless could give several instances various birds which have been known occasionally lay their eggs other birds nests now let suppose that the ancient progenitor our european cuckoo had the habits the american cuckoo but that occasionally she laid egg another bird nest the old bird profited this occasional habit the young were made more vigorous advantage having been taken the mistaken maternal instinct another bird than their own mother care encumbered she can hardly fail having eggs and young different ages the same time then the old birds the fostered young would gain advantage and analogy would lead believe that the young thus reared would apt follow inheritance the occasional and aberrant habit their mother and their turn would apt lay their eggs other birds nests and thus successful rearing their young continued process this nature believe that the strange instinct our cuckoo could and has been generated may add that according gray and some other observers the european cuckoo has not utterly lost all maternal love and care for her own offspring the occasional habit birds laying their eggs other birds nests either the same distinct species not very uncommon with the gallinaceæ and this perhaps explains the origin singular instinct the allied group ostriches for several hen ostriches least the case the american species unite and lay first few eggs one nest and then another and these are hatched the males this instinct may probably accounted for the fact the hens laying large number eggs but the case the cuckoo intervals two three days this instinct however the american ostrich has not yet been perfected for surprising number eggs lie strewed over the plains that one day hunting picked less than twenty lost and wasted eggs many bees are parasitic and always lay their eggs the nests bees other kinds this case more remarkable than that the cuckoo for these bees have not only their instincts but their structure modified accordance with their parasitic habits for they not possess the pollen collecting apparatus which would necessary they had store food for their own young some species likewise sphegidæ wasp like insects are parasitic other species and fabre has lately shown good reason for believing that although the tachytes nigra generally makes its own burrow and stores with paralysed prey for its own larvæ feed yet that when this insect finds burrow already made and stored another sphex takes advantage the prize and becomes for the occasion parasitic this case with the supposed case the cuckoo can see difficulty natural selection making occasional habit permanent advantage the species and the insect whose nest and stored food are thus feloniously appropriated not thus exterminated slave making instinct this remarkable instinct was first discovered the formica polyerges rufescens pierre huber better observer even than his celebrated father this ant absolutely dependent its slaves without their aid the species would certainly become extinct single year the males and fertile females work the workers sterile females though most energetic and courageous capturing slaves other work they are incapable making their own nests feeding their own larvæ when the old nest found inconvenient and they have migrate the slaves which determine the migration and actually carry their masters their jaws utterly helpless are the masters that when huber shut thirty them without slave but with plenty the food which they like best and with their larvæ and pupæ stimulate them work they did nothing they could not even feed themselves and many perished hunger huber then introduced single slave fusca and she instantly set work fed and saved the survivors made some cells and tended the larvæ and put all rights what can more extraordinary than these well ascertained facts had not known any other slave making ant would have been hopeless have speculated how wonderful instinct could have been perfected formica sanguinea was likewise first discovered huber slave making ant this species found the southern parts england and its habits have been attended smith the british museum whom much indebted for information this and other subjects although fully trusting the statements huber and smith tried approach the subject sceptical frame mind any one may well excused for doubting the truth extraordinary and odious instinct that making slaves hence will give the observations which have myself made some little detail opened fourteen nests sanguinea and found few slaves all males and fertile females the slave species are found only their own proper communities and have never been observed the nests sanguinea the slaves are black and not above half the size their red masters that the contrast their appearance very great when the nest slightly disturbed the slaves occasionally come out and like their masters are much agitated and defend the nest when the nest much disturbed and the larvæ and pupæ are exposed the slaves work energetically with their masters carrying them away place safety hence clear that the slaves feel quite home during the months june and july three successive years have watched for many hours several nests surrey and sussex and never saw slave either leave enter nest during these months the slaves are very few number thought that they might behave differently when more numerous but smith informs that has watched the nests various hours during may june and august both surrey and hampshire and has never seen the slaves though present large numbers august either leave enter the nest hence considers them strictly household slaves the masters the other hand may constantly seen bringing materials for the nest and food all kinds during the present year however the month july came across community with unusually large stock slaves and observed few slaves mingled with their masters leaving the nest and marching along the same road tall scotch fir tree twenty five yards distant which they ascended together probably search aphides cocci according huber who had ample opportunities for observation switzerland the slaves habitually work with their masters making the nest and they alone open and close the doors the morning and evening and huber expressly states their principal office search for aphides this difference the usual habits the masters and slaves the two countries probably depends merely the slaves being captured greater numbers switzerland than england one day fortunately chanced witness migration from one nest another and was most interesting spectacle behold the masters carefully carrying huber has described their slaves their jaws another day attention was struck about score the slave makers haunting the same spot and evidently not search food they approached and were vigorously repulsed independent community the slave species fusca sometimes many three these ants clinging the legs the slave making sanguinea the latter ruthlessly killed their small opponents and carried their dead bodies food their nest twenty nine yards distant but they were prevented from getting any pupæ rear slaves then dug small parcel the pupæ fusca from another nest and put them down bare spot near the place combat they were eagerly seized and carried off the tyrants who perhaps fancied that after all they had been victorious their late combat the same time laid the same place small parcel the pupæ another species flava with few these little yellow ants still clinging the fragments the nest this species sometimes though rarely made into slaves has been described smith although small species very courageous and have seen ferociously attack other ants one instance found surprise independent community flava under stone beneath nest the slave making sanguinea and when had accidentally disturbed both nests the little ants attacked their big neighbours with surprising courage now was curious ascertain whether sanguinea could distinguish the pupæ fusca which they habitually make into slaves from those the little and furious flava which they rarely capture and was evident that they did once distinguish them for have seen that they eagerly and instantly seized the pupæ fusca whereas they were much terrified when they came across the pupæ even the earth from the nest flava and quickly ran away but about quarter hour shortly after all the little yellow ants had crawled away they took heart and carried off the pupæ one evening visited another community sanguinea and found number these ants entering their nest carrying the dead bodies fusca showing that was not migration and numerous pupæ traced the returning file burthened with booty for about forty yards very thick clump heath whence saw the last individual sanguinea emerge carrying pupa but was not able find the desolated nest the thick heath the nest however must have been close hand for two three individuals fusca were rushing about the greatest agitation and one was perched motionless with its own pupa its mouth the top spray heath over its ravaged home such are the facts though they did not need confirmation regard the wonderful instinct making slaves let observed what contrast the instinctive habits sanguinea present with those the rufescens the latter does not build its own nest does not determine its own migrations does not collect food for itself its young and cannot even feed itself absolutely dependent its numerous slaves formica sanguinea the other hand possesses much fewer slaves and the early part the summer extremely few the masters determine when and where new nest shall formed and when they migrate the masters carry the slaves both switzerland and england the slaves seem have the exclusive care the larvæ and the masters alone slave making expeditions switzerland the slaves and masters work together making and bringing materials for the nest both but chiefly the slaves tend and milk may called their aphides and thus both collect food for the community england the masters alone usually leave the nest collect building materials and food for themselves their slaves and larvæ that the masters this country receive much less service from their slaves than they switzerland what steps the instinct sanguinea originated will not pretend conjecture but ants which are not slave makers will have seen carry off pupæ other species scattered near their nests possible that pupæ originally stored food might become developed and the ants thus unintentionally reared would then follow their proper instincts and what work they could their presence proved useful the species which had seized them were more advantageous this species capture workers than procreate them the habit collecting pupæ originally for food might natural selection strengthened and rendered permanent for the very different purpose raising slaves when the instinct was once acquired carried out much less extent even than our british sanguinea which have seen less aided its slaves than the same species switzerland can see difficulty natural selection increasing and modifying the instinct always supposing each modification use the species until ant was formed abjectly dependent its slaves the formica rufescens cell making instinct the hive bee will not here enter minute details this subject but will merely give outline the conclusions which have arrived must dull man who can examine the exquisite structure comb beautifully adapted its end without enthusiastic admiration hear from mathematicians that bees have practically solved recondite problem and have made their cells the proper shape hold the greatest possible amount honey with the least possible consumption precious wax their construction has been remarked that skilful workman with fitting tools and measures would find very difficult make cells wax the true form though this perfectly effected crowd bees working dark hive grant whatever instincts you please and seems first quite inconceivable how they can make all the necessary angles and planes even perceive when they are correctly made but the difficulty not nearly great first appears all this beautiful work can shown think follow from few very simple instincts was led investigate this subject waterhouse who has shown that the form the cell stands close relation the presence adjoining cells and the following view may perhaps considered only modification his theory let look the great principle gradation and see whether nature does not reveal her method work one end short series have humble bees which use their old cocoons hold honey sometimes adding them short tubes wax and likewise making separate and very irregular rounded cells wax the other end the series have the cells the hive bee placed double layer each cell well known hexagonal prism with the basal edges its six sides bevelled join pyramid formed three rhombs these rhombs have certain angles and the three which form the pyramidal base single cell one side the comb enter into the composition the bases three adjoining cells the opposite side the series between the extreme perfection the cells the hive bee and the simplicity those the humble bee have the cells the mexican melipona domestica carefully described and figured pierre huber the melipona itself intermediate structure between the hive and humble bee but more nearly related the latter forms nearly regular waxen comb cylindrical cells which the young are hatched and addition some large cells wax for holding honey these latter cells are nearly spherical and nearly equal sizes and are aggregated into irregular mass but the important point notice that these cells are always made that degree nearness each other that they would have intersected broken into each other the spheres had been completed but this never permitted the bees building perfectly flat walls wax between the spheres which thus tend intersect hence each cell consists outer spherical portion and two three more perfectly flat surfaces according the cell adjoins two three more other cells when one cell comes into contact with three other cells which from the spheres being nearly the same size very frequently and necessarily the case the three flat surfaces are united into pyramid and this pyramid huber has remarked manifestly gross imitation the three sided pyramidal basis the cell the hive bee the cells the hive bee here the three plane surfaces any one cell necessarily enter into the construction three adjoining cells obvious that the melipona saves wax this manner building for the flat walls between the adjoining cells are not double but are the same thickness the outer spherical portions and yet each flat portion forms part two cells reflecting this case occurred that the melipona had made its spheres some given distance from each other and had made them equal sizes and had arranged them symmetrically double layer the resulting structure would probably have been perfect the comb the hive bee accordingly wrote professor miller cambridge and this geometer has kindly read over the following statement drawn from his information and tells that strictly correct number equal spheres described with their centres placed two parallel layers with the centre each sphere the distance radius the square root radius 41421 some lesser distance from the centres the six surrounding spheres the same layer and the same distance from the centres the adjoining spheres the other and parallel layer then planes intersection between the several spheres both layers formed there will result double layer hexagonal prisms united together pyramidal bases formed three rhombs and the rhombs and the sides the hexagonal prisms will have every angle identically the same with the best measurements which have been made the cells the hive bee hence may safely conclude that could slightly modify the instincts already possessed the melipona and themselves not very wonderful this bee would make structure wonderfully perfect that the hive bee must suppose the melipona make her cells truly spherical and equal sizes and this would not very surprising seeing that she already does certain extent and seeing what perfectly cylindrical burrows wood many insects can make apparently turning round fixed point must suppose the melipona arrange her cells level layers she already does her cylindrical cells and must further suppose and this the greatest difficulty that she can somehow judge accurately what distance stand from her fellow labourers when several are making their spheres but she already far enabled judge distance that she always describes her spheres intersect largely and then she unites the points intersection perfectly flat surfaces have further suppose but this difficulty that after hexagonal prisms have been formed the intersection adjoining spheres the same layer she can prolong the hexagon any length requisite hold the stock honey the same way the rude humble bee adds cylinders wax the circular mouths her old cocoons such modifications instincts themselves not very wonderful hardly more wonderful than those which guide bird make its nest believe that the hive bee has acquired through natural selection her inimitable architectural powers but this theory can tested experiment following the example tegetmeier separated two combs and put between them long thick square strip wax the bees instantly began excavate minute circular pits and they deepened these little pits they made them wider and wider until they were converted into shallow basins appearing the eye perfectly true parts sphere and about the diameter cell was most interesting observe that wherever several bees had begun excavate these basins near together they had begun their work such distance from each other that the time the basins had acquired the above stated width about the width ordinary cell and were depth about one sixth the diameter the sphere which they formed part the rims the basins intersected broke into each other soon this occurred the bees ceased excavate and began build flat walls wax the lines intersection between the basins that each hexagonal prism was built upon the festooned edge smooth basin instead the straight edges three sided pyramid the case ordinary cells then put into the hive instead thick square piece wax thin and narrow knife edged ridge coloured with vermilion the bees instantly began both sides excavate little basins near each other the same way before but the ridge wax was thin that the bottoms the basins they had been excavated the same depth the former experiment would have broken into each other from the opposite sides the bees however did not suffer this happen and they stopped their excavations due time that the basins soon they had been little deepened came have flat bottoms and these flat bottoms formed thin little plates the vermilion wax having been left ungnawed were situated far the eye could judge exactly along the planes imaginary intersection between the basins the opposite sides the ridge wax parts only little bits other parts large portions rhombic plate had been left between the opposed basins but the work from the unnatural state things had not been neatly performed the bees must have worked very nearly the same rate the opposite sides the ridge vermilion wax they circularly gnawed away and deepened the basins both sides order have succeeded thus leaving flat plates between the basins stopping work along the intermediate planes planes intersection considering how flexible thin wax not see that there any difficulty the bees whilst work the two sides strip wax perceiving when they have gnawed the wax away the proper thinness and then stopping their work ordinary combs has appeared that the bees not always succeed working exactly the same rate from the opposite sides for have noticed half completed rhombs the base just commenced cell which were slightly concave one side where suppose that the bees had excavated too quickly and convex the opposed side where the bees had worked less quickly one well marked instance put the comb back into the hive and allowed the bees working for short time and again examined the cell and found that the rhombic plate had been completed and had become perfectly flat was absolutely impossible from the extreme thinness the little rhombic plate that they could have effected this gnawing away the convex side and suspect that the bees such cases stand the opposed cells and push and bend the ductile and warm wax which have tried easily done into its proper intermediate plane and thus flatten from the experiment the ridge vermilion wax can clearly see that the bees were build for themselves thin wall wax they could make their cells the proper shape standing the proper distance from each other excavating the same rate and endeavouring make equal spherical hollows but never allowing the spheres break into each other now bees may clearly seen examining the edge growing comb make rough circumferential wall rim all round the comb and they gnaw into this from the opposite sides always working circularly they deepen each cell they not make the whole three sided pyramidal base any one cell the same time but only the one rhombic plate which stands the extreme growing margin the two plates the case may and they never complete the upper edges the rhombic plates until the hexagonal walls are commenced some these statements differ from those made the justly celebrated elder huber but convinced their accuracy and had space could show that they are conformable with theory huber statement that the very first cell excavated out little parallel sided wall wax not far have seen strictly correct the first commencement having always been little hood wax but will not here enter these details see how important part excavation plays the construction the cells but would great error suppose that the bees cannot build rough wall wax the proper position that along the plane intersection between two adjoining spheres have several specimens showing clearly that they can this even the rude circumferential rim wall wax round growing comb flexures may sometimes observed corresponding position the planes the rhombic basal plates future cells but the rough wall wax has every case finished off being largely gnawed away both sides the manner which the bees build curious they always make the first rough wall from ten twenty times thicker than the excessively thin finished wall the cell which will ultimately left shall understand how they work supposing masons first pile broad ridge cement and then begin cutting away equally both sides near the ground till smooth very thin wall left the middle the masons always piling the cut away cement and adding fresh cement the summit the ridge shall thus have thin wall steadily growing upward but always crowned gigantic coping from all the cells both those just commenced and those completed being thus crowned strong coping wax the bees can cluster and crawl over the comb without injuring the delicate hexagonal walls which are only about one four hundredth inch thickness the plates the pyramidal basis being about twice thick this singular manner building strength continually given the comb with the utmost ultimate economy wax seems first add the difficulty understanding how the cells are made that multitude bees all work together one bee after working short time one cell going another that huber has stated score individuals work even the commencement the first cell was able practically show this fact covering the edges the hexagonal walls single cell the extreme margin the circumferential rim growing comb with extremely thin layer melted vermilion wax and invariably found that the colour was most delicately diffused the bees delicately painter could have done with his brush atoms the coloured wax having been taken from the spot which had been placed and worked into the growing edges the cells all round the work construction seems sort balance struck between many bees all instinctively standing the same relative distance from each other all trying sweep equal spheres and then building leaving ungnawed the planes intersection between these spheres was really curious note cases difficulty when two pieces comb met angle how often the bees would entirely pull down and rebuild different ways the same cell sometimes recurring shape which they had first rejected when bees have place which they can stand their proper positions for working for instance slip wood placed directly under the middle comb growing downwards that the comb has built over one face the slip this case the bees can lay the foundations one wall new hexagon its strictly proper place projecting beyond the other completed cells suffices that the bees should enabled stand their proper relative distances from each other and from the walls the last completed cells and then striking imaginary spheres they can build wall intermediate between two adjoining spheres but far have seen they never gnaw away and finish off the angles cell till large part both that cell and the adjoining cells has been built this capacity bees laying down under certain circumstances rough wall its proper place between two just commenced cells important bears fact which seems first quite subversive the foregoing theory namely that the cells the extreme margin wasp combs are sometimes strictly hexagonal but have not space here enter this subject nor does there seem any great difficulty single insect the case queen wasp making hexagonal cells she work alternately the inside and outside two three cells commenced the same time always standing the proper relative distance from the parts the cells just begun sweeping spheres cylinders and building intermediate planes even conceivable that insect might fixing point which commence cell and then moving outside first one point and then five other points the proper relative distances from the central point and from each other strike the planes intersection and make isolated hexagon but not aware that any such case has been observed nor would any good derived from single hexagon being built its construction more materials would required than for cylinder natural selection acts only the accumulation slight modifications structure instinct each profitable the individual under its conditions life may reasonably asked how long and graduated succession modified architectural instincts all tending towards the present perfect plan construction could have profited the progenitors the hive bee think the answer not difficult known that bees are often hard pressed get sufficient nectar and informed tegetmeier that has been experimentally found that less than from twelve fifteen pounds dry sugar are consumed hive bees for the secretion each pound wax that prodigious quantity fluid nectar must collected and consumed the bees hive for the secretion the wax necessary for the construction their combs moreover many bees have remain idle for many days during the process secretion large store honey indispensable support large stock bees during the winter and the security the hive known mainly depend large number bees being supported hence the saving wax largely saving honey must most important element success any family bees course the success any species bee may dependent the number its parasites other enemies quite distinct causes and altogether independent the quantity honey which the bees could collect but let suppose that this latter circumstance determined probably often does determine the numbers humble bee which could exist country and let further suppose that the community lived throughout the winter and consequently required store honey there can this case doubt that would advantage our humble bee slight modification her instinct led her make her waxen cells near together intersect little for wall common even two adjoining cells would save some little wax hence would continually more and more advantageous our humble bee she were make her cells more and more regular nearer together and aggregated into mass like the cells the melipona for this case large part the bounding surface each cell would serve bound other cells and much wax would saved again from the same cause would advantageous the melipona she were make her cells closer together and more regular every way than present for then have seen the spherical surfaces would wholly disappear and would all replaced plane surfaces and the melipona would make comb perfect that the hive bee beyond this stage perfection architecture natural selection could not lead for the comb the hive bee far can see absolutely perfect economising wax thus believe the most wonderful all known instincts that the hive bee can explained natural selection having taken advantage numerous successive slight modifications simpler instincts natural selection having slow degrees more and more perfectly led the bees sweep equal spheres given distance from each other double layer and build and excavate the wax along the planes intersection the bees course more knowing that they swept their spheres one particular distance from each other than they know what are the several angles the hexagonal prisms and the basal rhombic plates the motive power the process natural selection having been economy wax that individual swarm which wasted least honey the secretion wax having succeeded best and having transmitted inheritance its newly acquired economical instinct new swarms which their turn will have had the best chance succeeding the struggle for existence doubt many instincts very difficult explanation could opposed the theory natural selection cases which cannot see how instinct could possibly have originated cases which intermediate gradations are known exist cases instinct apparently such trifling importance that they could hardly have been acted natural selection cases instincts almost identically the same animals remote the scale nature that cannot account for their similarity inheritance from common parent and must therefore believe that they have been acquired independent acts natural selection will not here enter these several cases but will confine myself one special difficulty which first appeared insuperable and actually fatal whole theory allude the neuters sterile females insect communities for these neuters often differ widely instinct and structure from both the males and fertile females and yet from being sterile they cannot propagate their kind the subject well deserves discussed great length but will here take only single case that working sterile ants how the workers have been rendered sterile difficulty but not much greater than that any other striking modification structure for can shown that some insects and other articulate animals state nature occasionally become sterile and such insects had been social and had been profitable the community that number should have been annually born capable work but incapable procreation can see very great difficulty this being effected natural selection but must pass over this preliminary difficulty the great difficulty lies the working ants differing widely from both the males and the fertile females structure the shape the thorax and being destitute wings and sometimes eyes and instinct far instinct alone concerned the prodigious difference this respect between the workers and the perfect females would have been far better exemplified the hive bee working ant other neuter insect had been animal the ordinary state should have unhesitatingly assumed that all its characters had been slowly acquired through natural selection namely individual having been born with some slight profitable modification structure this being inherited its offspring which again varied and were again selected and onwards but with the working ant have insect differing greatly from its parents yet absolutely sterile that could never have transmitted successively acquired modifications structure instinct its progeny may well asked how possible reconcile this case with the theory natural selection first let remembered that have innumerable instances both our domestic productions and those state nature all sorts differences structure which have become correlated certain ages and either sex have differences correlated not only one sex but that short period alone when the reproductive system active the nuptial plumage many birds and the hooked jaws the male salmon have even slight differences the horns different breeds cattle relation artificially imperfect state the male sex for oxen certain breeds have longer horns than other breeds comparison with the horns the bulls cows these same breeds hence can see real difficulty any character having become correlated with the sterile condition certain members insect communities the difficulty lies understanding how such correlated modifications structure could have been slowly accumulated natural selection this difficulty though appearing insuperable lessened believe disappears when remembered that selection may applied the family well the individual and may thus gain the desired end thus well flavoured vegetable cooked and the individual destroyed but the horticulturist sows seeds the same stock and confidently expects get nearly the same variety breeders cattle wish the flesh and fat well marbled together the animal has been slaughtered but the breeder goes with confidence the same family have such faith the powers selection that not doubt that breed cattle always yielding oxen with extraordinarily long horns could slowly formed carefully watching which individual bulls and cows when matched produced oxen with the longest horns and yet one could ever have propagated its kind thus believe has been with social insects slight modification structure instinct correlated with the sterile condition certain members the community has been advantageous the community consequently the fertile males and females the same community flourished and transmitted their fertile offspring tendency produce sterile members having the same modification and believe that this process has been repeated until that prodigious amount difference between the fertile and sterile females the same species has been produced which see many social insects but have not yet touched the climax the difficulty namely the fact that the neuters several ants differ not only from the fertile females and males but from each other sometimes almost incredible degree and are thus divided into two even three castes the castes moreover not generally graduate into each other but are perfectly well defined being distinct from each other are any two species the same genus rather any two genera the same family thus eciton there are working and soldier neuters with jaws and instincts extraordinarily different cryptocerus the workers one caste alone carry wonderful sort shield their heads the use which quite unknown the mexican myrmecocystus the workers one caste never leave the nest they are fed the workers another caste and they have enormously developed abdomen which secretes sort honey supplying the place that excreted the aphides the domestic cattle they may called which our european ants guard imprison will indeed thought that have overweening confidence the principle natural selection when not admit that such wonderful and well established facts once annihilate theory the simpler case neuter insects all one caste the same kind which have been rendered natural selection believe quite possible different from the fertile males and females this case may safely conclude from the analogy ordinary variations that each successive slight profitable modification did not probably first appear all the individual neuters the same nest but few alone and that the long continued selection the fertile parents which produced most neuters with the profitable modification all the neuters ultimately came have the desired character this view ought occasionally find neuter insects the same species the same nest presenting gradations structure and this find even often considering how few neuter insects out europe have been carefully examined smith has shown how surprisingly the neuters several british ants differ from each other size and sometimes colour and that the extreme forms can sometimes perfectly linked together individuals taken out the same nest have myself compared perfect gradations this kind often happens that the larger the smaller sized workers are the most numerous that both large and small are numerous with those intermediate size scanty numbers formica flava has larger and smaller workers with some intermediate size and this species smith has observed the larger workers have simple eyes ocelli which though small can plainly distinguished whereas the smaller workers have their ocelli rudimentary having carefully dissected several specimens these workers can affirm that the eyes are far more rudimentary the smaller workers than can accounted for merely their proportionally lesser size and fully believe though dare not assert positively that the workers intermediate size have their ocelli exactly intermediate condition that here have two bodies sterile workers the same nest differing not only size but their organs vision yet connected some few members intermediate condition may digress adding that the smaller workers had been the most useful the community and those males and females had been continually selected which produced more and more the smaller workers until all the workers had come this condition should then have had species ant with neuters very nearly the same condition with those myrmica for the workers myrmica have not even rudiments ocelli though the male and female ants this genus have well developed ocelli may give one other case confidently did expect find gradations important points structure between the different castes neuters the same species that gladly availed myself smith offer numerous specimens from the same nest the driver ant anomma west africa the reader will perhaps best appreciate the amount difference these workers giving not the actual measurements but strictly accurate illustration the difference was the same were see set workmen building house whom many were five feet four inches high and many sixteen feet high but must suppose that the larger workmen had heads four instead three times big those the smaller men and jaws nearly five times big the jaws moreover the working ants the several sizes differed wonderfully shape and the form and number the teeth but the important fact for that though the workers can grouped into castes different sizes yet they graduate insensibly into each other does the widely different structure their jaws speak confidently this latter point lubbock made drawings for with the camera lucida the jaws which had dissected from the workers the several sizes with these facts before believe that natural selection acting the fertile parents could form species which should regularly produce neuters either all large size with one form jaw all small size with jaws having widely different structure lastly and this our climax difficulty one set workers one size and structure and simultaneously another set workers different size and structure graduated series having been first formed the case the driver ant and then the extreme forms from being the most useful the community having been produced greater and greater numbers through the natural selection the parents which generated them until none with intermediate structure were produced thus believe the wonderful fact two distinctly defined castes sterile workers existing the same nest both widely different from each other and from their parents has originated can see how useful their production may have been social community insects the same principle that the division labour useful civilised man ants work inherited instincts and inherited tools weapons and not acquired knowledge and manufactured instruments perfect division labour could effected with them only the workers being sterile for had they been fertile they would have intercrossed and their instincts and structure would have become blended and nature has believe effected this admirable division labour the communities ants the means natural selection but bound confess that with all faith this principle should never have anticipated that natural selection could have been efficient high degree had not the case these neuter insects convinced the fact have therefore discussed this case some little but wholly insufficient length order show the power natural selection and likewise because this far the most serious special difficulty which theory has encountered the case also very interesting proves that with animals with plants any amount modification structure can effected the accumulation numerous slight and must call them accidental variations which are any manner profitable without exercise habit having come into play for amount exercise habit volition the utterly sterile members community could possibly have affected the structure instincts the fertile members which alone leave descendants surprised that one has advanced this demonstrative case neuter insects against the well known doctrine lamarck summary have endeavoured briefly this chapter show that the mental qualities our domestic animals vary and that the variations are inherited still more briefly have attempted show that instincts vary slightly state nature one will dispute that instincts are the highest importance each animal therefore can see difficulty under changing conditions life natural selection accumulating slight modifications instinct any extent any useful direction some cases habit use and disuse have probably come into play not pretend that the facts given this chapter strengthen any great degree theory but none the cases difficulty the best judgment annihilate the other hand the fact that instincts are not always absolutely perfect and are liable mistakes that instinct has been produced for the exclusive good other animals but that each animal takes advantage the instincts others that the canon natural history natura non facit saltum applicable instincts well corporeal structure and plainly explicable the foregoing views but otherwise inexplicable all tend corroborate the theory natural selection this theory also strengthened some few other facts regard instincts that common case closely allied but certainly distinct species when inhabiting distant parts the world and living under considerably different conditions life yet often retaining nearly the same instincts for instance can understand the principle inheritance how that the thrush south america lines its nest with mud the same peculiar manner does our british thrush how that the male wrens troglodytes north america build cock nests roost like the males our distinct kitty wrens habit wholly unlike that any other known bird finally may not logical deduction but imagination far more satisfactory look such instincts the young cuckoo ejecting its foster brothers ants making slaves the larvæ ichneumonidæ feeding within the live bodies caterpillars not specially endowed created instincts but small consequences one general law leading the advancement all organic beings namely multiply vary let the strongest live and the weakest die chapter viii hybridism distinction between the sterility first crosses and hybrids sterility various degree not universal affected close interbreeding removed domestication laws governing the sterility hybrids sterility not special endowment but incidental other differences causes the sterility first crosses and hybrids parallelism between the effects changed conditions life and crossing fertility varieties when crossed and their mongrel offspring not universal hybrids and mongrels compared independently their fertility summary the view generally entertained naturalists that species when intercrossed have been specially endowed with the quality sterility order prevent the confusion all organic forms this view certainly seems first probable for species within the same country could hardly have kept distinct had they been capable crossing freely the importance the fact that hybrids are very generally sterile has think been much underrated some late writers the theory natural selection the case especially important inasmuch the sterility hybrids could not possibly any advantage them and therefore could not have been acquired the continued preservation successive profitable degrees sterility hope however able show that sterility not specially acquired endowed quality but incidental other acquired differences treating this subject two classes facts large extent fundamentally different have generally been confounded together namely the sterility two species when first crossed and the sterility the hybrids produced from them pure species have course their organs reproduction perfect condition yet when intercrossed they produce either few offspring hybrids the other hand have their reproductive organs functionally impotent may clearly seen the state the male element both plants and animals though the organs themselves are perfect structure far the microscope reveals the first case the two sexual elements which form the embryo are perfect the second case they are either not all developed are imperfectly developed this distinction important when the cause the sterility which common the two cases has considered the distinction has probably been slurred over owing the sterility both cases being looked special endowment beyond the province our reasoning powers the fertility varieties that the forms known believed have descended from common parents when intercrossed and likewise the fertility their mongrel offspring theory equal importance with the sterility species for seems make broad and clear distinction between varieties and species first for the sterility species when crossed and their hybrid offspring impossible study the several memoirs and works those two conscientious and admirable observers kölreuter and gärtner who almost devoted their lives this subject without being deeply impressed with the high generality some degree sterility kölreuter makes the rule universal but then cuts the knot for ten cases which found two forms considered most authors distinct species quite fertile together unhesitatingly ranks them varieties gärtner also makes the rule equally universal and disputes the entire fertility kölreuter ten cases but these and many other cases gärtner obliged carefully count the seeds order show that there any degree sterility always compares the maximum number seeds produced two species when crossed and their hybrid offspring with the average number produced both pure parent species state nature but serious cause error seems here introduced plant hybridised must castrated and what often more important must secluded order prevent pollen being brought insects from other plants nearly all the plants experimentised gärtner were potted and apparently were kept chamber his house that these processes are often injurious the fertility plant cannot doubted for gärtner gives his table about score cases plants which castrated and artificially fertilised with their own pollen and excluding all cases such the leguminosæ which there acknowledged difficulty the manipulation half these twenty plants had their fertility some degree impaired moreover gärtner during several years repeatedly crossed the primrose and cowslip which have such good reason believe varieties and only once twice succeeded getting fertile seed found the common red and blue pimpernels anagallis arvensis and coerulea which the best botanists rank varieties absolutely sterile together and came the same conclusion several other analogous cases seems that may well permitted doubt whether many other species are really sterile when intercrossed gärtner believes certain the one hand that the sterility various species when crossed different degree and graduates away insensibly and the other hand that the fertility pure species easily affected various circumstances that for all practical purposes most difficult say where perfect fertility ends and sterility begins think better evidence this can required than that the two most experienced observers who have ever lived namely kölreuter and gärtner should have arrived diametrically opposite conclusions regard the very same species also most instructive compare but have not space here enter details the evidence advanced our best botanists the question whether certain doubtful forms should ranked species varieties with the evidence from fertility adduced different hybridisers the same author from experiments made during different years can thus shown that neither sterility nor fertility affords any clear distinction between species and varieties but that the evidence from this source graduates away and doubtful the same degree the evidence derived from other constitutional and structural differences regard the sterility hybrids successive generations though gärtner was enabled rear some hybrids carefully guarding them from cross with either pure parent for six seven and one case for ten generations yet asserts positively that their fertility never increased but generally greatly decreased not doubt that this usually the case and that the fertility often suddenly decreases the first few generations nevertheless believe that all these experiments the fertility has been diminished independent cause namely from close interbreeding have collected large body facts showing that close interbreeding lessens fertility and the other hand that occasional cross with distinct individual variety increases fertility that cannot doubt the correctness this almost universal belief amongst breeders hybrids are seldom raised experimentalists great numbers and the parent species other allied hybrids generally grow the same garden the visits insects must carefully prevented during the flowering season hence hybrids will generally fertilised during each generation their own individual pollen and convinced that this would injurious their fertility already lessened their hybrid origin strengthened this conviction remarkable statement repeatedly made gärtner namely that even the less fertile hybrids artificially fertilised with hybrid pollen the same kind their fertility notwithstanding the frequent ill effects manipulation sometimes decidedly increases and goes increasing now artificial fertilisation pollen often taken chance know from own experience from the anthers another flower from the anthers the flower itself which fertilised that cross between two flowers though probably the same plant would thus effected moreover whenever complicated experiments are progress careful observer gärtner would have castrated his hybrids and this would have insured each generation cross with the pollen from distinct flower either from the same plant from another plant the same hybrid nature and thus the strange fact the increase fertility the successive generations artificially fertilised hybrids may believe accounted for close interbreeding having been avoided now let turn the results arrived the third most experienced hybridiser namely the honourable and reverend herbert emphatic his conclusion that some hybrids are perfectly fertile fertile the pure parent species are kölreuter and gärtner that some degree sterility between distinct species universal law nature experimentised some the very same species did gärtner the difference their results may think part accounted for herbert great horticultural skill and his having hothouses his command his many important statements will here give only single one example namely that every ovule pod crinum capense fertilised revolutum produced plant which says never saw occur case its natural fecundation that here have perfect even more than commonly perfect fertility first cross between two distinct species this case the crinum leads refer most singular fact namely that there are individual plants with certain species lobelia and with all the species the genus hippeastrum which can far more easily fertilised the pollen another and distinct species than their own pollen for these plants have been found yield seed the pollen distinct species though quite sterile with their own pollen notwithstanding that their own pollen was found perfectly good for fertilised distinct species that certain individual plants and all the individuals certain species can actually hybridised much more readily than they can self fertilised for instance bulb hippeastrum aulicum produced four flowers three were fertilised herbert with their own pollen and the fourth was subsequently fertilised the pollen compound hybrid descended from three other and distinct species the result was that the ovaries the three first flowers soon ceased grow and after few days perished entirely whereas the pod impregnated the pollen the hybrid made vigorous growth and rapid progress maturity and bore good seed which vegetated freely letter 1839 herbert told that had then tried the experiment during five years and continued try during several subsequent years and always with the same result this result has also been confirmed other observers the case hippeastrum with its sub genera and the case some other genera lobelia passiflora and verbascum although the plants these experiments appeared perfectly healthy and although both the ovules and pollen the same flower were perfectly good with respect other species yet they were functionally imperfect their mutual self action must infer that the plants were unnatural state nevertheless these facts show what slight and mysterious causes the lesser greater fertility species when crossed comparison with the same species when self fertilised sometimes depends the practical experiments horticulturists though not made with scientific precision deserve some notice notorious how complicated manner the species pelargonium fuchsia calceolaria petunia rhododendron etc have been crossed yet many these hybrids seed freely for instance herbert asserts that hybrid from calceolaria integrifolia and plantaginea species most widely dissimilar general habit reproduced itself perfectly had been natural species from the mountains chile have taken some pains ascertain the degree fertility some the complex crosses rhododendrons and assured that many them are perfectly fertile noble for instance informs that raises stocks for grafting from hybrid between rhododendron ponticum and catawbiense and that this hybrid seeds freely possible imagine had hybrids when fairly treated gone decreasing fertility each successive generation gärtner believes the case the fact would have been notorious nurserymen horticulturists raise large beds the same hybrids and such alone are fairly treated for insect agency the several individuals the same hybrid variety are allowed freely cross with each other and the injurious influence close interbreeding thus prevented any one may readily convince himself the efficiency insect agency examining the flowers the more sterile kinds hybrid rhododendrons which produce pollen for will find their stigmas plenty pollen brought from other flowers regard animals much fewer experiments have been carefully tried than with plants our systematic arrangements can trusted that the genera animals are distinct from each other are the genera plants then may infer that animals more widely separated the scale nature can more easily crossed than the case plants but the hybrids themselves are think more sterile doubt whether any case perfectly fertile hybrid animal can considered thoroughly well authenticated should however borne mind that owing few animals breeding freely under confinement few experiments have been fairly tried for instance the canary bird has been crossed with nine other finches but not one these nine species breeds freely confinement have right expect that the first crosses between them and the canary that their hybrids should perfectly fertile again with respect the fertility successive generations the more fertile hybrid animals hardly know instance which two families the same hybrid have been raised the same time from different parents avoid the ill effects close interbreeding the contrary brothers and sisters have usually been crossed each successive generation opposition the constantly repeated admonition every breeder and this case not all surprising that the inherent sterility the hybrids should have gone increasing were act thus and pair brothers and sisters the case any pure animal which from any cause had the least tendency sterility the breed would assuredly lost very few generations although not know any thoroughly well authenticated cases perfectly fertile hybrid animals have some reason believe that the hybrids from cervulus vaginalis and reevesii and from phasianus colchicus with torquatus and with versicolor are perfectly fertile the hybrids from the common and chinese geese cygnoides species which are different that they are generally ranked distinct genera have often bred this country with either pure parent and one single instance they have bred inter this was effected eyton who raised two hybrids from the same parents but from different hatches and from these two birds raised less than eight hybrids grandchildren the pure geese from one nest india however these cross bred geese must far more fertile for assured two eminently capable judges namely blyth and capt hutton that whole flocks these crossed geese are kept various parts the country and they are kept for profit where neither pure parent species exists they must certainly highly fertile doctrine which originated with pallas has been largely accepted modern naturalists namely that most our domestic animals have descended from two more aboriginal species since commingled intercrossing this view the aboriginal species must either first have produced quite fertile hybrids the hybrids must have become subsequent generations quite fertile under domestication this latter alternative seems the most probable and inclined believe its truth although rests direct evidence believe for instance that our dogs have descended from several wild stocks yet with perhaps the exception certain indigenous domestic dogs south america all are quite fertile together and analogy makes greatly doubt whether the several aboriginal species would first have freely bred together and have produced quite fertile hybrids again there reason believe that our european and the humped indian cattle are quite fertile together but from facts communicated blyth think they must considered distinct species this view the origin many our domestic animals must either give the belief the almost universal sterility distinct species animals when crossed must look sterility not indelible characteristic but one capable being removed domestication finally looking all the ascertained facts the intercrossing plants and animals may concluded that some degree sterility both first crosses and hybrids extremely general result but that cannot under our present state knowledge considered absolutely universal laws governing the sterility first crosses and hybrids will now consider little more detail the circumstances and rules governing the sterility first crosses and hybrids our chief object will see whether not the rules indicate that species have specially been endowed with this quality order prevent their crossing and blending together utter confusion the following rules and conclusions are chiefly drawn from gärtner admirable work the hybridisation plants have taken much pains ascertain how far the rules apply animals and considering how scanty our knowledge regard hybrid animals have been surprised find how generally the same rules apply both kingdoms has been already remarked that the degree fertility both first crosses and hybrids graduates from zero perfect fertility surprising how many curious ways this gradation can shown exist but only the barest outline the facts can here given when pollen from plant one family placed the stigma plant distinct family exerts more influence than much inorganic dust from this absolute zero fertility the pollen different species the same genus applied the stigma some one species yields perfect gradation the number seeds produced nearly complete even quite complete fertility and have seen certain abnormal cases even excess fertility beyond that which the plant own pollen will produce hybrids themselves there are some which never have produced and probably never would produce even with the pollen either pure parent single fertile seed but some these cases first trace fertility may detected the pollen one the pure parent species causing the flower the hybrid wither earlier than otherwise would have done and the early withering the flower well known sign incipient fertilisation from this extreme degree sterility have self fertilised hybrids producing greater and greater number seeds perfect fertility hybrids from two species which are very difficult cross and which rarely produce any offspring are generally very sterile but the parallelism between the difficulty making first cross and the sterility the hybrids thus produced two classes facts which are generally confounded together means strict there are many cases which two pure species can united with unusual facility and produce numerous hybrid offspring yet these hybrids are remarkably sterile the other hand there are species which can crossed very rarely with extreme difficulty but the hybrids when last produced are very fertile even within the limits the same genus for instance dianthus these two opposite cases occur the fertility both first crosses and hybrids more easily affected unfavourable conditions than the fertility pure species but the degree fertility likewise innately variable for not always the same when the same two species are crossed under the same circumstances but depends part upon the constitution the individuals which happen have been chosen for the experiment with hybrids for their degree fertility often found differ greatly the several individuals raised from seed out the same capsule and exposed exactly the same conditions the term systematic affinity meant the resemblance between species structure and constitution more especially the structure parts which are high physiological importance and which differ little the allied species now the fertility first crosses between species and the hybrids produced from them largely governed their systematic affinity this clearly shown hybrids never having been raised between species ranked systematists distinct families and the other hand very closely allied species generally uniting with facility but the correspondence between systematic affinity and the facility crossing means strict multitude cases could given very closely allied species which will not unite only with extreme difficulty and the other hand very distinct species which unite with the utmost facility the same family there may genus dianthus which very many species can most readily crossed and another genus silene which the most persevering efforts have failed produce between extremely close species single hybrid even within the limits the same genus meet with this same difference for instance the many species nicotiana have been more largely crossed than the species almost any other genus but gärtner found that acuminata which not particularly distinct species obstinately failed fertilise fertilised less than eight other species nicotiana very many analogous facts could given one has been able point out what kind what amount difference any recognisable character sufficient prevent two species crossing can shown that plants most widely different habit and general appearance and having strongly marked differences every part the flower even the pollen the fruit and the cotyledons can crossed annual and perennial plants deciduous and evergreen trees plants inhabiting different stations and fitted for extremely different climates can often crossed with ease reciprocal cross between two species mean the case for instance stallion horse being first crossed with female ass and then male ass with mare these two species may then said have been reciprocally crossed there often the widest possible difference the facility making reciprocal crosses such cases are highly important for they prove that the capacity any two species cross often completely independent their systematic affinity any recognisable difference their whole organisation the other hand these cases clearly show that the capacity for crossing connected with constitutional differences imperceptible and confined the reproductive system this difference the result reciprocal crosses between the same two species was long ago observed kölreuter give instance mirabilis jalappa can easily fertilised the pollen longiflora and the hybrids thus produced are sufficiently fertile but kölreuter tried more than two hundred times during eight following years fertilise reciprocally longiflora with the pollen jalappa and utterly failed several other equally striking cases could given thuret has observed the same fact with certain sea weeds fuci gärtner moreover found that this difference facility making reciprocal crosses extremely common lesser degree has observed even between forms closely related matthiola annua and glabra that many botanists rank them only varieties also remarkable fact that hybrids raised from reciprocal crosses though course compounded the very same two species the one species having first been used the father and then the mother generally differ fertility small and occasionally high degree several other singular rules could given from gärtner for instance some species have remarkable power crossing with other species other species the same genus have remarkable power impressing their likeness their hybrid offspring but these two powers not all necessarily together there are certain hybrids which instead having usual intermediate character between their two parents always closely resemble one them and such hybrids though externally like one their pure parent species are with rare exceptions extremely sterile again amongst hybrids which are usually intermediate structure between their parents exceptional and abnormal individuals sometimes are born which closely resemble one their pure parents and these hybrids are almost always utterly sterile even when the other hybrids raised from seed from the same capsule have considerable degree fertility these facts show how completely fertility the hybrid independent its external resemblance either pure parent considering the several rules now given which govern the fertility first crosses and hybrids see that when forms which must considered good and distinct species are united their fertility graduates from zero perfect fertility even fertility under certain conditions excess that their fertility besides being eminently susceptible favourable and unfavourable conditions innately variable that means always the same degree the first cross and the hybrids produced from this cross that the fertility hybrids not related the degree which they resemble external appearance either parent and lastly that the facility making first cross between any two species not always governed their systematic affinity degree resemblance each other this latter statement clearly proved reciprocal crosses between the same two species for according the one species the other used the father the mother there generally some difference and occasionally the widest possible difference the facility effecting union the hybrids moreover produced from reciprocal crosses often differ fertility now these complex and singular rules indicate that species have been endowed with sterility simply prevent their becoming confounded nature think not for why should the sterility extremely different degree when various species are crossed all which must suppose would equally important keep from blending together why should the degree sterility innately variable the individuals the same species why should some species cross with facility and yet produce very sterile hybrids and other species cross with extreme difficulty and yet produce fairly fertile hybrids why should there often great difference the result reciprocal cross between the same two species why may even asked has the production hybrids been permitted grant species the special power producing hybrids and then stop their further propagation different degrees sterility not strictly related the facility the first union between their parents seems strange arrangement the foregoing rules and facts the other hand appear clearly indicate that the sterility both first crosses and hybrids simply incidental dependent unknown differences chiefly the reproductive systems the species which are crossed the differences being peculiar and limited nature that reciprocal crosses between two species the male sexual element the one will often freely act the female sexual element the other but not reversed direction will advisable explain little more fully example what mean sterility being incidental other differences and not specially endowed quality the capacity one plant grafted budded another entirely unimportant for its welfare state nature presume that one will suppose that this capacity specially endowed quality but will admit that incidental differences the laws growth the two plants can sometimes see the reason why one tree will not take another from differences their rate growth the hardness their wood the period the flow nature their sap etc but multitude cases can assign reason whatever great diversity the size two plants one being woody and the other herbaceous one being evergreen and the other deciduous and adaptation widely different climates does not always prevent the two grafting together hybridisation with grafting the capacity limited systematic affinity for one has been able graft trees together belonging quite distinct families and the other hand closely allied species and varieties the same species can usually but not invariably grafted with ease but this capacity hybridisation means absolutely governed systematic affinity although many distinct genera within the same family have been grafted together other cases species the same genus will not take each other the pear can grafted far more readily the quince which ranked distinct genus than the apple which member the same genus even different varieties the pear take with different degrees facility the quince different varieties the apricot and peach certain varieties the plum gärtner found that there was sometimes innate difference different individuals the same two species crossing sagaret believes this the case with different individuals the same two species being grafted together reciprocal crosses the facility effecting union often very far from equal sometimes grafting the common gooseberry for instance cannot grafted the currant whereas the currant will take though with difficulty the gooseberry have seen that the sterility hybrids which have their reproductive organs imperfect condition very different case from the difficulty uniting two pure species which have their reproductive organs perfect yet these two distinct cases run certain extent parallel something analogous occurs grafting for thouin found that three species robinia which seeded freely their own roots and which could grafted with great difficulty another species when thus grafted were rendered barren the other hand certain species sorbus when grafted other species yielded twice much fruit when their own roots are reminded this latter fact the extraordinary case hippeastrum lobelia etc which seeded much more freely when fertilised with the pollen distinct species than when self fertilised with their own pollen thus see that although there clear and fundamental difference between the mere adhesion grafted stocks and the union the male and female elements the act reproduction yet that there rude degree parallelism the results grafting and crossing distinct species and must look the curious and complex laws governing the facility with which trees can grafted each other incidental unknown differences their vegetative systems believe that the still more complex laws governing the facility first crosses are incidental unknown differences chiefly their reproductive systems these differences both cases follow certain extent might have been expected systematic affinity which every kind resemblance and dissimilarity between organic beings attempted expressed the facts means seem indicate that the greater lesser difficulty either grafting crossing together various species has been special endowment although the case crossing the difficulty important for the endurance and stability specific forms the case grafting unimportant for their welfare causes the sterility first crosses and hybrids may now look little closer the probable causes the sterility first crosses and hybrids these two cases are fundamentally different for just remarked the union two pure species the male and female sexual elements are perfect whereas hybrids they are imperfect even first crosses the greater lesser difficulty effecting union apparently depends several distinct causes there must sometimes physical impossibility the male element reaching the ovule would the case with plant having pistil too long for the pollen tubes reach the ovarium has also been observed that when pollen one species placed the stigma distantly allied species though the pollen tubes protrude they not penetrate the stigmatic surface again the male element may reach the female element but incapable causing embryo developed seems have been the case with some thuret experiments fuci explanation can given these facts any more than why certain trees cannot grafted others lastly embryo may developed and then perish early period this latter alternative has not been sufficiently attended but believe from observations communicated hewitt who has had great experience hybridising gallinaceous birds that the early death the embryo very frequent cause sterility first crosses was first very unwilling believe this view hybrids when once born are generally healthy and long lived see the case the common mule hybrids however are differently circumstanced before and after birth when born and living country where their two parents can live they are generally placed under suitable conditions life but hybrid partakes only half the nature and constitution its mother and therefore before birth long nourished within its mother womb within the egg seed produced the mother may exposed conditions some degree unsuitable and consequently liable perish early period more especially all very young beings seem eminently sensitive injurious unnatural conditions life regard the sterility hybrids which the sexual elements are imperfectly developed the case very different have more than once alluded large body facts which have collected showing that when animals and plants are removed from their natural conditions they are extremely liable have their reproductive systems seriously affected this fact the great bar the domestication animals between the sterility thus superinduced and that hybrids there are many points similarity both cases the sterility independent general health and often accompanied excess size great luxuriance both cases the sterility occurs various degrees both the male element the most liable affected but sometimes the female more than the male both the tendency goes certain extent with systematic affinity for whole groups animals and plants are rendered impotent the same unnatural conditions and whole groups species tend produce sterile hybrids the other hand one species group will sometimes resist great changes conditions with unimpaired fertility and certain species group will produce unusually fertile hybrids one can tell till tries whether any particular animal will breed under confinement any plant seed freely under culture nor can tell till tries whether any two species genus will produce more less sterile hybrids lastly when organic beings are placed during several generations under conditions not natural them they are extremely liable vary which due believe their reproductive systems having been specially affected though lesser degree than when sterility ensues with hybrids for hybrids successive generations are eminently liable vary every experimentalist has observed thus see that when organic beings are placed under new and unnatural conditions and when hybrids are produced the unnatural crossing two species the reproductive system independently the general state health affected sterility very similar manner the one case the conditions life have been disturbed though often slight degree inappreciable the other case that hybrids the external conditions have remained the same but the organisation has been disturbed two different structures and constitutions having been blended into one for scarcely possible that two organisations should compounded into one without some disturbance occurring the development periodical action mutual relation the different parts and organs one another the conditions life when hybrids are able breed inter they transmit their offspring from generation generation the same compounded organisation and hence need not surprised that their sterility though some degree variable rarely diminishes must however confessed that cannot understand excepting vague hypotheses several facts with respect the sterility hybrids for instance the unequal fertility hybrids produced from reciprocal crosses the increased sterility those hybrids which occasionally and exceptionally resemble closely either pure parent nor pretend that the foregoing remarks the root the matter explanation offered why organism when placed under unnatural conditions rendered sterile all that have attempted show that two cases some respects allied sterility the common result the one case from the conditions life having been disturbed the other case from the organisation having been disturbed two organisations having been compounded into one may seem fanciful but suspect that similar parallelism extends allied yet very different class facts old and almost universal belief founded think considerable body evidence that slight changes the conditions life are beneficial all living things see this acted farmers and gardeners their frequent exchanges seed tubers etc from one soil climate another and back again during the convalescence animals plainly see that great benefit derived from almost any change the habits life again both with plants and animals there abundant evidence that cross between very distinct individuals the same species that between members different strains sub breeds gives vigour and fertility the offspring believe indeed from the facts alluded our fourth chapter that certain amount crossing indispensable even with hermaphrodites and that close interbreeding continued during several generations between the nearest relations especially these kept under the same conditions life always induces weakness and sterility the progeny hence seems that the one hand slight changes the conditions life benefit all organic beings and the other hand that slight crosses that crosses between the males and females the same species which have varied and become slightly different give vigour and fertility the offspring but have seen that greater changes changes particular nature often render organic beings some degree sterile and that greater crosses that crosses between males and females which have become widely specifically different produce hybrids which are generally sterile some degree cannot persuade myself that this parallelism accident illusion both series facts seem connected together some common but unknown bond which essentially related the principle life fertility varieties when crossed and their mongrel offspring may urged most forcible argument that there must some essential distinction between species and varieties and that there must some error all the foregoing remarks inasmuch varieties however much they may differ from each other external appearance cross with perfect facility and yield perfectly fertile offspring fully admit that this almost invariably the case but look varieties produced under nature are immediately involved hopeless difficulties for two hitherto reputed varieties found any degree sterile together they are once ranked most naturalists species for instance the blue and red pimpernel the primrose and cowslip which are considered many our best botanists varieties are said gärtner not quite fertile when crossed and consequently ranks them undoubted species thus argue circle the fertility all varieties produced under nature will assuredly have granted turn varieties produced supposed have been produced under domestication are still involved doubt for when stated for instance that the german spitz dog unites more easily than other dogs with foxes that certain south american indigenous domestic dogs not readily cross with european dogs the explanation which will occur everyone and probably the true one that these dogs have descended from several aboriginally distinct species nevertheless the perfect fertility many domestic varieties differing widely from each other appearance for instance the pigeon the cabbage remarkable fact more especially when reflect how many species there are which though resembling each other most closely are utterly sterile when intercrossed several considerations however render the fertility domestic varieties less remarkable than first appears can the first place clearly shown that mere external dissimilarity between two species does not determine their greater lesser degree sterility when crossed and may apply the same rule domestic varieties the second place some eminent naturalists believe that long course domestication tends eliminate sterility the successive generations hybrids which were first only slightly sterile and this surely ought not expect find sterility both appearing and disappearing under nearly the same conditions life lastly and this seems far the most important consideration new races animals and plants are produced under domestication man methodical and unconscious power selection for his own use and pleasure neither wishes select nor could select slight differences the reproductive system other constitutional differences correlated with the reproductive system supplies his several varieties with the same food treats them nearly the same manner and does not wish alter their general habits life nature acts uniformly and slowly during vast periods time the whole organisation any way which may for each creature own good and thus she may either directly more probably indirectly through correlation modify the reproductive system the several descendants from any one species seeing this difference the process selection carried man and nature need not surprised some difference the result have yet spoken the varieties the same species were invariably fertile when intercrossed but seems impossible resist the evidence the existence certain amount sterility the few following cases which will briefly abstract the evidence least good that from which believe the sterility multitude species the evidence also derived from hostile witnesses who all other cases consider fertility and sterility safe criterions specific distinction gärtner kept during several years dwarf kind maize with yellow seeds and tall variety with red seeds growing near each other his garden and although these plants have separated sexes they never naturally crossed then fertilised thirteen flowers the one with the pollen the other but only single head produced any seed and this one head produced only five grains manipulation this case could not have been injurious the plants have separated sexes one believe has suspected that these varieties maize are distinct species and important notice that the hybrid plants thus raised were themselves perfectly fertile that even gärtner did not venture consider the two varieties specifically distinct girou buzareingues crossed three varieties gourd which like the maize has separated sexes and asserts that their mutual fertilisation much the less easy their differences are greater how far these experiments may trusted know not but the forms experimentised are ranked sagaret who mainly founds his classification the test infertility varieties the following case far more remarkable and seems first quite incredible but the result astonishing number experiments made during many years nine species verbascum good observer and hostile witness gärtner namely that yellow and white varieties the same species verbascum when intercrossed produce less seed than either coloured varieties when fertilised with pollen from their own coloured flowers moreover asserts that when yellow and white varieties one species are crossed with yellow and white varieties distinct species more seed produced the crosses between the same coloured flowers than between those which are differently coloured yet these varieties verbascum present other difference besides the mere colour the flower and one variety can sometimes raised from the seed the other from observations which have made certain varieties hollyhock inclined suspect that they present analogous facts kölreuter whose accuracy has been confirmed every subsequent observer has proved the remarkable fact that one variety the common tobacco more fertile when crossed with widely distinct species than are the other varieties experimentised five forms which are commonly reputed varieties and which tested the severest trial namely reciprocal crosses and found their mongrel offspring perfectly fertile but one these five varieties when used either father mother and crossed with the nicotiana glutinosa always yielded hybrids not sterile those which were produced from the four other varieties when crossed with glutinosa hence the reproductive system this one variety must have been some manner and some degree modified from these facts from the great difficulty ascertaining the infertility varieties state nature for supposed variety infertile any degree would generally ranked species from man selecting only external characters the production the most distinct domestic varieties and from not wishing being able produce recondite and functional differences the reproductive system from these several considerations and facts not think that the very general fertility varieties can proved universal occurrence form fundamental distinction between varieties and species the general fertility varieties does not seem sufficient overthrow the view which have taken with respect the very general but not invariable sterility first crosses and hybrids namely that not special endowment but incidental slowly acquired modifications more especially the reproductive systems the forms which are crossed hybrids and mongrels compared independently their fertility independently the question fertility the offspring species when crossed and varieties when crossed may compared several other respects gärtner whose strong wish was draw marked line distinction between species and varieties could find very few and seems quite unimportant differences between the called hybrid offspring species and the called mongrel offspring varieties and the other hand they agree most closely very many important respects shall here discuss this subject with extreme brevity the most important distinction that the first generation mongrels are more variable than hybrids but gärtner admits that hybrids from species which have long been cultivated are often variable the first generation and have myself seen striking instances this fact gärtner further admits that hybrids between very closely allied species are more variable than those from very distinct species and this shows that the difference the degree variability graduates away when mongrels and the more fertile hybrids are propagated for several generations extreme amount variability their offspring notorious but some few cases both hybrids and mongrels long retaining uniformity character could given the variability however the successive generations mongrels perhaps greater than hybrids this greater variability mongrels than hybrids does not seem all surprising for the parents mongrels are varieties and mostly domestic varieties very few experiments having been tried natural varieties and this implies most cases that there has been recent variability and therefore might expect that such variability would often continue and super added that arising from the mere act crossing the slight degree variability hybrids from the first cross the first generation contrast with their extreme variability the succeeding generations curious fact and deserves attention for bears and corroborates the view which have taken the cause ordinary variability namely that due the reproductive system being eminently sensitive any change the conditions life being thus often rendered either impotent least incapable its proper function producing offspring identical with the parent form now hybrids the first generation are descended from species excluding those long cultivated which have not had their reproductive systems any way affected and they are not variable but hybrids themselves have their reproductive systems seriously affected and their descendants are highly variable but return our comparison mongrels and hybrids gärtner states that mongrels are more liable than hybrids revert either parent form but this true certainly only difference degree gärtner further insists that when any two species although most closely allied each other are crossed with third species the hybrids are widely different from each other whereas two very distinct varieties one species are crossed with another species the hybrids not differ much but this conclusion far can make out founded single experiment and seems directly opposed the results several experiments made kölreuter these alone are the unimportant differences which gärtner able point out between hybrid and mongrel plants the other hand the resemblance mongrels and hybrids their respective parents more especially hybrids produced from nearly related species follows according gärtner the same laws when two species are crossed one has sometimes prepotent power impressing its likeness the hybrid and believe with varieties plants with animals one variety certainly often has this prepotent power over another variety hybrid plants produced from reciprocal cross generally resemble each other closely and with mongrels from reciprocal cross both hybrids and mongrels can reduced either pure parent form repeated crosses successive generations with either parent these several remarks are apparently applicable animals but the subject here excessively complicated partly owing the existence secondary sexual characters but more especially owing prepotency transmitting likeness running more strongly one sex than the other both when one species crossed with another and when one variety crossed with another variety for instance think those authors are right who maintain that the ass has prepotent power over the horse that both the mule and the hinny more resemble the ass than the horse but that the prepotency runs more strongly the male ass than the female that the mule which the offspring the male ass and mare more like ass than the hinny which the offspring the female ass and stallion much stress has been laid some authors the supposed fact that mongrel animals alone are born closely like one their parents but can shown that this does sometimes occur with hybrids yet grant much less frequently with hybrids than with mongrels looking the cases which have collected cross bred animals closely resembling one parent the resemblances seem chiefly confined characters almost monstrous their nature and which have suddenly appeared such albinism melanism deficiency tail horns additional fingers and toes and not relate characters which have been slowly acquired selection consequently sudden reversions the perfect character either parent would more likely occur with mongrels which are descended from varieties often suddenly produced and semi monstrous character than with hybrids which are descended from species slowly and naturally produced the whole entirely agree with prosper lucas who after arranging enormous body facts with respect animals comes the conclusion that the laws resemblance the child its parents are the same whether the two parents differ much little from each other namely the union individuals the same variety different varieties distinct species laying aside the question fertility and sterility all other respects there seems general and close similarity the offspring crossed species and crossed varieties look species having been specially created and varieties having been produced secondary laws this similarity would astonishing fact but harmonises perfectly with the view that there essential distinction between species and varieties summary chapter first crosses between forms sufficiently distinct ranked species and their hybrids are very generally but not universally sterile the sterility all degrees and often slight that the two most careful experimentalists who have ever lived have come diametrically opposite conclusions ranking forms this test the sterility innately variable individuals the same species and eminently susceptible favourable and unfavourable conditions the degree sterility does not strictly follow systematic affinity but governed several curious and complex laws generally different and sometimes widely different reciprocal crosses between the same two species not always equal degree first cross and the hybrid produced from this cross the same manner grafting trees the capacity one species variety take another incidental generally unknown differences their vegetative systems crossing the greater less facility one species unite with another incidental unknown differences their reproductive systems there more reason think that species have been specially endowed with various degrees sterility prevent them crossing and blending nature than think that trees have been specially endowed with various and somewhat analogous degrees difficulty being grafted together order prevent them becoming inarched our forests the sterility first crosses between pure species which have their reproductive systems perfect seems depend several circumstances some cases largely the early death the embryo the sterility hybrids which have their reproductive systems imperfect and which have had this system and their whole organisation disturbed being compounded two distinct species seems closely allied that sterility which frequently affects pure species when their natural conditions life have been disturbed this view supported parallelism another kind namely that the crossing forms only slightly different favourable the vigour and fertility their offspring and that slight changes the conditions life are apparently favourable the vigour and fertility all organic beings not surprising that the degree difficulty uniting two species and the degree sterility their hybrid offspring should generally correspond though due distinct causes for both depend the amount difference some kind between the species which are crossed nor surprising that the facility effecting first cross the fertility the hybrids produced and the capacity being grafted together though this latter capacity evidently depends widely different circumstances should all run certain extent parallel with the systematic affinity the forms which are subjected experiment for systematic affinity attempts express all kinds resemblance between all species first crosses between forms known varieties sufficiently alike considered varieties and their mongrel offspring are very generally but not quite universally fertile nor this nearly general and perfect fertility surprising when remember how liable are argue circle with respect varieties state nature and when remember that the greater number varieties have been produced under domestication the selection mere external differences and not differences the reproductive system all other respects excluding fertility there close general resemblance between hybrids and mongrels finally then the facts briefly given this chapter not seem opposed but even rather support the view that there fundamental distinction between species and varieties chapter the imperfection the geological record the absence intermediate varieties the present day the nature extinct intermediate varieties their number the vast lapse time inferred from the rate deposition and denudation the poorness our palæontological collections the intermittence geological formations the absence intermediate varieties any one formation the sudden appearance groups species their sudden appearance the lowest known fossiliferous strata the sixth chapter enumerated the chief objections which might justly urged against the views maintained this volume most them have now been discussed one namely the distinctness specific forms and their not being blended together innumerable transitional links very obvious difficulty assigned reasons why such links not commonly occur the present day under the circumstances apparently most favourable for their presence namely extensive and continuous area with graduated physical conditions endeavoured show that the life each species depends more important manner the presence other already defined organic forms than climate and therefore that the really governing conditions life not graduate away quite insensibly like heat moisture endeavoured also show that intermediate varieties from existing lesser numbers than the forms which they connect will generally beaten out and exterminated during the course further modification and improvement the main cause however innumerable intermediate links not now occurring everywhere throughout nature depends the very process natural selection through which new varieties continually take the places and exterminate their parent forms but just proportion this process extermination has acted enormous scale must the number intermediate varieties which have formerly existed the earth truly enormous why then not every geological formation and every stratum full such intermediate links geology assuredly does not reveal any such finely graduated organic chain and this perhaps the most obvious and gravest objection which can urged against theory the explanation lies believe the extreme imperfection the geological record the first place should always borne mind what sort intermediate forms must theory have formerly existed have found difficult when looking any two species avoid picturing myself forms directly intermediate between them but this wholly false view should always look for forms intermediate between each species and common but unknown progenitor and the progenitor will generally have differed some respects from all its modified descendants give simple illustration the fantail and pouter pigeons have both descended from the rock pigeon possessed all the intermediate varieties which have ever existed should have extremely close series between both and the rock pigeon but should have varieties directly intermediate between the fantail and pouter none for instance combining tail somewhat expanded with crop somewhat enlarged the characteristic features these two breeds these two breeds moreover have become much modified that had historical indirect evidence regarding their origin would not have been possible have determined from mere comparison their structure with that the rock pigeon whether they had descended from this species from some other allied species such oenas with natural species look forms very distinct for instance the horse and tapir have reason suppose that links ever existed directly intermediate between them but between each and unknown common parent the common parent will have had its whole organisation much general resemblance the tapir and the horse but some points structure may have differed considerably from both even perhaps more than they differ from each other hence all such cases should unable recognise the parent form any two more species even closely compared the structure the parent with that its modified descendants unless the same time had nearly perfect chain the intermediate links just possible theory that one two living forms might have descended from the other for instance horse from tapir and this case direct intermediate links will have existed between them but such case would imply that one form had remained for very long period unaltered whilst its descendants had undergone vast amount change and the principle competition between organism and organism between child and parent will render this very rare event for all cases the new and improved forms life will tend supplant the old and unimproved forms the theory natural selection all living species have been connected with the parent species each genus differences not greater than see between the varieties the same species the present day and these parent species now generally extinct have their turn been similarly connected with more ancient species and backwards always converging the common ancestor each great class that the number intermediate and transitional links between all living and extinct species must have been inconceivably great but assuredly this theory true such have lived upon this earth the lapse time independently our not finding fossil remains such infinitely numerous connecting links may objected that time will not have sufficed for great amount organic change all changes having been effected very slowly through natural selection hardly possible for even recall the reader who may not practical geologist the facts leading the mind feebly comprehend the lapse time who can read sir charles lyell grand work the principles geology which the future historian will recognise having produced revolution natural science yet does not admit how incomprehensibly vast have been the past periods time may once close this volume not that suffices study the principles geology read special treatises different observers separate formations and mark how each author attempts give inadequate idea the duration each formation even each stratum man must for years examine for himself great piles superimposed strata and watch the sea work grinding down old rocks and making fresh sediment before can hope comprehend anything the lapse time the monuments which see around good wander along lines sea coast when formed moderately hard rocks and mark the process degradation the tides most cases reach the cliffs only for short time twice day and the waves eat into them only when they are charged with sand pebbles for there reason believe that pure water can effect little nothing wearing away rock last the base the cliff undermined huge fragments fall down and these remaining fixed have worn away atom atom until reduced size they can rolled about the waves and then are more quickly ground into pebbles sand mud but how often see along the bases retreating cliffs rounded boulders all thickly clothed marine productions showing how little they are abraded and how seldom they are rolled about moreover follow for few miles any line rocky cliff which undergoing degradation find that only here and there along short length round promontory that the cliffs are the present time suffering the appearance the surface and the vegetation show that elsewhere years have elapsed since the waters washed their base who most closely studies the action the sea our shores will believe most deeply impressed with the slowness with which rocky coasts are worn away the observations this head hugh miller and that excellent observer smith jordan hill are most impressive with the mind thus impressed let any one examine beds conglomerate many thousand feet thickness which though probably formed quicker rate than many other deposits yet from being formed worn and rounded pebbles each which bears the stamp time are good show how slowly the mass has been accumulated let him remember lyell profound remark that the thickness and extent sedimentary formations are the result and measure the degradation which the earth crust has elsewhere suffered and what amount degradation implied the sedimentary deposits many countries professor ramsay has given the maximum thickness most cases from actual measurement few cases from estimate each formation different parts great britain and this the result feet palæozoic strata not including igneous beds 154 secondary strata 190 tertiary strata 240 making altogether 584 feet that very nearly thirteen and three quarters british miles some these formations which are represented england thin beds are thousands feet thickness the continent moreover between each successive formation have the opinion most geologists enormously long blank periods that the lofty pile sedimentary rocks britain gives but inadequate idea the time which has elapsed during their accumulation yet what time this must have consumed good observers have estimated that sediment deposited the great mississippi river the rate only 600 feet hundred thousand years this estimate may quite erroneous yet considering over what wide spaces very fine sediment transported the currents the sea the process accumulation any one area must extremely slow but the amount denudation which the strata have many places suffered independently the rate accumulation the degraded matter probably offers the best evidence the lapse time remember having been much struck with the evidence denudation when viewing volcanic islands which have been worn the waves and pared all round into perpendicular cliffs one two thousand feet height for the gentle slope the lava streams due their formerly liquid state showed glance how far the hard rocky beds had once extended into the open ocean the same story still more plainly told faults those great cracks along which the strata have been upheaved one side thrown down the other the height depth thousands feet for since the crust cracked the surface the land has been completely planed down the action the sea that trace these vast dislocations externally visible the craven fault for instance extends for upwards miles and along this line the vertical displacement the strata has varied from 600 3000 feet professor ramsay has published account downthrow anglesea 2300 feet and informs that fully believes there one merionethshire 000 feet yet these cases there nothing the surface show such prodigious movements the pile rocks the one other side having been smoothly swept away the consideration these facts impresses mind almost the same manner does the vain endeavour grapple with the idea eternity tempted give one other case the well known one the denudation the weald though must admitted that the denudation the weald has been mere trifle comparison with that which has removed masses our palæozoic strata parts ten thousand feet thickness shown professor ramsay masterly memoir this subject yet admirable lesson stand the north downs and look the distant south downs for remembering that great distance the west the northern and southern escarpments meet and close one can safely picture oneself the great dome rocks which must have covered the weald within limited period since the latter part the chalk formation the distance from the northern the southern downs about miles and the thickness the several formations average about 1100 feet informed professor ramsay but some geologists suppose range older rocks underlies the weald the flanks which the overlying sedimentary deposits might have accumulated thinner masses than elsewhere the above estimate would erroneous but this source doubt probably would not greatly affect the estimate applied the western extremity the district then knew the rate which the sea commonly wears away line cliff any given height could measure the time requisite have denuded the weald this course cannot done but may order form some crude notion the subject assume that the sea would eat into cliffs 500 feet height the rate one inch century this will first appear much too small allowance but the same were assume cliff one yard height eaten back along whole line coast the rate one yard nearly every twenty two years doubt whether any rock even soft chalk would yield this rate excepting the most exposed coasts though doubt the degradation lofty cliff would more rapid from the breakage the fallen fragments the other hand not believe that any line coast ten twenty miles length ever suffers degradation the same time along its whole indented length and must remember that almost all strata contain harder layers nodules which from long resisting attrition form breakwater the base hence under ordinary circumstances conclude that for cliff 500 feet height denudation one inch per century for the whole length would ample allowance this rate the above data the denudation the weald must have required 306 662 400 years say three hundred million years the action fresh water the gently inclined wealden district when upraised could hardly have been great but would somewhat reduce the above estimate the other hand during oscillations level which know this area has undergone the surface may have existed for millions years land and thus have escaped the action the sea when deeply submerged for perhaps equally long periods would likewise have escaped the action the coast waves that all probability far longer period than 300 million years has elapsed since the latter part the secondary period have made these few remarks because highly important for gain some notion however imperfect the lapse years during each these years over the whole world the land and the water has been peopled hosts living forms what infinite number generations which the mind cannot grasp must have succeeded each other the long roll years now turn our richest geological museums and what paltry display behold the poorness our palæontological collections that our palæontological collections are very imperfect admitted every one the remark that admirable palæontologist the late edward forbes should not forgotten namely that numbers our fossil species are known and named from single and often broken specimens from few specimens collected some one spot only small portion the surface the earth has been geologically explored and part with sufficient care the important discoveries made every year europe prove organism wholly soft can preserved shells and bones will decay and disappear when left the bottom the sea where sediment not accumulating believe are continually taking most erroneous view when tacitly admit ourselves that sediment being deposited over nearly the whole bed the sea rate sufficiently quick embed and preserve fossil remains throughout enormously large proportion the ocean the bright blue tint the water bespeaks its purity the many cases record formation conformably covered after enormous interval time another and later formation without the underlying bed having suffered the interval any wear and tear seem explicable only the view the bottom the sea not rarely lying for ages unaltered condition the remains which become embedded sand gravel will when the beds are upraised generally dissolved the percolation rain water suspect that but few the very many animals which live the beach between high and low watermark are preserved for instance the several species the chthamalinæ sub family sessile cirripedes coat the rocks all over the world infinite numbers they are all strictly littoral with the exception single mediterranean species which inhabits deep water and has been found fossil sicily whereas not one other species has hitherto been found any tertiary formation yet now known that the genus chthamalus existed during the chalk period the molluscan genus chiton offers partially analogous case with respect the terrestrial productions which lived during the secondary and palæozoic periods superfluous state that our evidence from fossil remains fragmentary extreme degree for instance not land shell known belonging either these vast periods with one exception discovered sir lyell the carboniferous strata north america regard mammiferous remains single glance the historical table published the supplement lyell manual will bring home the truth how accidental and rare their preservation far better than pages detail nor their rarity surprising when remember how large proportion the bones tertiary mammals have been discovered either caves lacustrine deposits and that not cave true lacustrine bed known belonging the age our secondary palæozoic formations but the imperfection the geological record mainly results from another and more important cause than any the foregoing namely from the several formations being separated from each other wide intervals time when see the formations tabulated written works when follow them nature difficult avoid believing that they are closely consecutive but know for instance from sir murchison great work russia what wide gaps there are that country between the superimposed formations north america and many other parts the world the most skilful geologist his attention had been exclusively confined these large territories would never have suspected that during the periods which were blank and barren his own country great piles sediment charged with new and peculiar forms life had elsewhere been accumulated and each separate territory hardly any idea can formed the length time which has elapsed between the consecutive formations may infer that this could nowhere ascertained the frequent and great changes the mineralogical composition consecutive formations generally implying great changes the geography the surrounding lands whence the sediment has been derived accords with the belief vast intervals time having elapsed between each formation but can think see why the geological formations each region are almost invariably intermittent that have not followed each other close sequence scarcely any fact struck more when examining many hundred miles the south american coasts which have been upraised several hundred feet within the recent period than the absence any recent deposits sufficiently extensive last for even short geological period along the whole west coast which inhabited peculiar marine fauna tertiary beds are scantily developed that record several successive and peculiar marine faunas will probably preserved distant age little reflection will explain why along the rising coast the western side south america extensive formations with recent tertiary remains can anywhere found though the supply sediment must for ages have been great from the enormous degradation the coast rocks and from muddy streams entering the sea the explanation doubt that the littoral and sub littoral deposits are continually worn away soon they are brought the slow and gradual rising the land within the grinding action the coast waves may think safely conclude that sediment must accumulated extremely thick solid extensive masses order withstand the incessant action the waves when first upraised and during subsequent oscillations level such thick and extensive accumulations sediment may formed two ways either profound depths the sea which case judging from the researches forbes may conclude that the bottom will inhabited extremely few animals and the mass when upraised will give most imperfect record the forms life which then existed sediment may accumulated any thickness and extent over shallow bottom continue slowly subside this latter case long the rate subsidence and supply sediment nearly balance each other the sea will remain shallow and favourable for life and thus fossiliferous formation thick enough when upraised resist any amount degradation may formed convinced that all our ancient formations which are rich fossils have thus been formed during subsidence since publishing views this subject 1845 have watched the progress geology and have been surprised note how author after author treating this that great formation has come the conclusion that was accumulated during subsidence may add that the only ancient tertiary formation the west coast south america which has been bulky enough resist such degradation has yet suffered but which will hardly last distant geological age was certainly deposited during downward oscillation level and thus gained considerable thickness all geological facts tell plainly that each area has undergone numerous slow oscillations level and apparently these oscillations have affected wide spaces consequently formations rich fossils and sufficiently thick and extensive resist subsequent degradation may have been formed over wide spaces during periods subsidence but only where the supply sediment was sufficient keep the sea shallow and embed and preserve the remains before they had time decay the other hand long the bed the sea remained stationary thick deposits could not have been accumulated the shallow parts which are the most favourable life still less could this have happened during the alternate periods elevation speak more accurately the beds which were then accumulated will have been destroyed being upraised and brought within the limits the coast action thus the geological record will almost necessarily rendered intermittent feel much confidence the truth these views for they are strict accordance with the general principles inculcated sir lyell and forbes independently arrived similar conclusion one remark here worth passing notice during periods elevation the area the land and the adjoining shoal parts the sea will increased and new stations will often formed all circumstances most favourable previously explained for the formation new varieties and species but during such periods there will generally blank the geological record the other hand during subsidence the inhabited area and number inhabitants will decrease excepting the productions the shores continent when first broken into archipelago and consequently during subsidence though there will much extinction fewer new varieties species will formed and during these very periods subsidence that our great deposits rich fossils have been accumulated nature may almost said have guarded against the frequent discovery her transitional linking forms from the foregoing considerations cannot doubted that the geological record viewed whole extremely imperfect but confine our attention any one formation becomes more difficult understand why not therein find closely graduated varieties between the allied species which lived its commencement and its close some cases are record the same species presenting distinct varieties the upper and lower parts the same formation but they are rare they may here passed over although each formation has indisputably required vast number years for its deposition can see several reasons why each should not include graduated series links between the species which then lived but can means pretend assign due proportional weight the following considerations although each formation may mark very long lapse years each perhaps short compared with the period requisite change one species into another aware that two palæontologists whose opinions are worthy much deference namely bronn and woodward have concluded that the average duration each formation twice thrice long the average duration specific forms but insuperable difficulties seems prevent coming any just conclusion this head when see species first appearing the middle any formation would rash the extreme infer that had not elsewhere previously existed again when find species disappearing before the uppermost layers have been deposited would equally rash suppose that then became wholly extinct forget how small the area europe compared with the rest the world nor have the several stages the same formation throughout europe been correlated with perfect accuracy with marine animals all kinds may safely infer large amount migration during climatal and other changes and when see species first appearing any formation the probability that only then first immigrated into that area well known for instance that several species appeared somewhat earlier the palæozoic beds north america than those europe time having apparently been required for their migration from the american the european seas examining the latest deposits various quarters the world has everywhere been noted that some few still existing species are common the deposit but have become extinct the immediately surrounding sea conversely that some are now abundant the neighbouring sea but are rare absent this particular deposit excellent lesson reflect the ascertained amount migration the inhabitants europe during the glacial period which forms only part one whole geological period and likewise reflect the great changes level the inordinately great change climate the prodigious lapse time all included within this same glacial period yet may doubted whether any quarter the world sedimentary deposits including fossil remains have gone accumulating within the same area during the whole this period not for instance probable that sediment was deposited during the whole the glacial period near the mouth the mississippi within that limit depth which marine animals can flourish for know what vast geographical changes occurred other parts america during this space time when such beds were deposited shallow water near the mouth the mississippi during some part the glacial period shall have been upraised organic remains will probably first appear and disappear different levels owing the migration species and geographical changes and the distant future geologist examining these beds might tempted conclude that the average duration life the embedded fossils had been less than that the glacial period instead having been really far greater that extending from before the glacial epoch the present day order get perfect gradation between two forms the upper and lower parts the same formation the deposit must have gone accumulating for very long period order have given sufficient time for the slow process variation hence the deposit will generally have very thick one and the species undergoing modification will have had live the same area throughout this whole time but have seen that thick fossiliferous formation can only accumulated during period subsidence and keep the depth approximately the same which necessary order enable the same species live the same space the supply sediment must nearly have counterbalanced the amount subsidence but this same movement subsidence will often tend sink the area whence the sediment derived and thus diminish the supply whilst the downward movement continues fact this nearly exact balancing between the supply sediment and the amount subsidence probably rare contingency for has been observed more than one palæontologist that very thick deposits are usually barren organic remains except near their upper lower limits would seem that each separate formation like the whole pile formations any country has generally been intermittent its accumulation when see often the case formation composed beds different mineralogical composition may reasonably suspect that the process deposition has been much interrupted change the currents the sea and supply sediment different nature will generally have been due geographical changes requiring much time nor will the closest inspection formation give any idea the time which its deposition has consumed many instances could given beds only few feet thickness representing formations elsewhere thousands feet thickness and which must have required enormous period for their accumulation yet one ignorant this fact would have suspected the vast lapse time represented the thinner formation many cases could given the lower beds formation having been upraised denuded submerged and then covered the upper beds the same formation facts showing what wide yet easily overlooked intervals have occurred its accumulation other cases have the plainest evidence great fossilised trees still standing upright they grew many long intervals time and changes level during the process deposition which would never even have been suspected had not the trees chanced have been preserved thus messrs lyell and dawson found carboniferous beds 1400 feet thick nova scotia with ancient root bearing strata one above the other less than sixty eight different levels hence when the same species occur the bottom middle and top formation the probability that they have not lived the same spot during the whole period deposition but have disappeared and reappeared perhaps many times during the same geological period that such species were undergo considerable amount modification during any one geological period section would not probably include all the fine intermediate gradations which must theory have existed between them but abrupt though perhaps very slight changes form all important remember that naturalists have golden rule which distinguish species and varieties they grant some little variability each species but when they meet with somewhat greater amount difference between any two forms they rank both species unless they are enabled connect them together close intermediate gradations and this from the reasons just assigned can seldom hope effect any one geological section supposing and two species and third found underlying bed even were strictly intermediate between and would simply ranked third and distinct species unless the same time could most closely connected with either one both forms intermediate varieties nor should forgotten before explained that might the actual progenitor and and yet might not all necessarily strictly intermediate between them all points structure that might obtain the parent species and its several modified descendants from the lower and upper beds formation and unless obtained numerous transitional gradations should not recognise their relationship and should consequently compelled rank them all distinct species notorious what excessively slight differences many palæontologists have founded their species and they this the more readily the specimens come from different sub stages the same formation some experienced conchologists are now sinking many the very fine species orbigny and others into the rank varieties and this view find the kind evidence change which theory ought find moreover look rather wider intervals namely distinct but consecutive stages the same great formation find that the embedded fossils though almost universally ranked specifically different yet are far more closely allied each other than are the species found more widely separated formations but this subject shall have return the following chapter one other consideration worth notice with animals and plants that can propagate rapidly and are not highly locomotive there reason suspect have formerly seen that their varieties are generally first local and that such local varieties not spread widely and supplant their parent forms until they have been modified and perfected some considerable degree according this view the chance discovering formation any one country all the early stages transition between any two forms small for the successive changes are supposed have been local confined some one spot most marine animals have wide range and have seen that with plants those which have the widest range that oftenest present varieties that with shells and other marine animals probably those which have had the widest range far exceeding the limits the known geological formations europe which have oftenest given rise first local varieties and ultimately new species and this again would greatly lessen the chance our being able trace the stages transition any one geological formation should not forgotten that the present day with perfect specimens for examination two forms can seldom connected intermediate varieties and thus proved the same species until many specimens have been collected from many places and the case fossil species this could rarely effected palæontologists shall perhaps best perceive the improbability our being enabled connect species numerous fine intermediate fossil links asking ourselves whether for instance geologists some future period will able prove that our different breeds cattle sheep horses and dogs have descended from single stock from several aboriginal stocks again whether certain sea shells inhabiting the shores north america which are ranked some conchologists distinct species from their european representatives and other conchologists only varieties are really varieties are called specifically distinct this could effected only the future geologist discovering fossil state numerous intermediate gradations and such success seems improbable the highest degree geological research though has added numerous species existing and extinct genera and has made the intervals between some few groups less wide than they otherwise would have been yet has done scarcely anything breaking down the distinction between species connecting them together numerous fine intermediate varieties and this not having been effected probably the gravest and most obvious all the many objections which may urged against views hence will worth while sum the foregoing remarks under imaginary illustration the malay archipelago about the size europe from the north cape the mediterranean and from britain russia and therefore equals all the geological formations which have been examined with any accuracy excepting those the united states america fully agree with godwin austen that the present condition the malay archipelago with its numerous large islands separated wide and shallow seas probably represents the former state europe when most our formations were accumulating the malay archipelago one the richest regions the whole world organic beings yet all the species were collected which have ever lived there how imperfectly would they represent the natural history the world but have every reason believe that the terrestrial productions the archipelago would preserved excessively imperfect manner the formations which suppose there accumulating suspect that not many the strictly littoral animals those which lived naked submarine rocks would embedded and those embedded gravel sand would not endure distant epoch wherever sediment did not accumulate the bed the sea where did not accumulate sufficient rate protect organic bodies from decay remains could preserved our archipelago believe that fossiliferous formations could formed sufficient thickness last age distant futurity the secondary formations lie the past only during periods subsidence these periods subsidence would separated from each other enormous intervals during which the area would either stationary rising whilst rising each fossiliferous formation would destroyed almost soon accumulated the incessant coast action now see the shores south america during the periods subsidence there would probably much extinction life during the periods elevation there would much variation but the geological record would then least perfect may doubted whether the duration any one great period subsidence over the whole part the archipelago together with contemporaneous accumulation sediment would exceed the average duration the same specific forms and these contingencies are indispensable for the preservation all the transitional gradations between any two more species such gradations were not fully preserved transitional varieties would merely appear many distinct species also probable that each great period subsidence would interrupted oscillations level and that slight climatal changes would intervene during such lengthy periods and these cases the inhabitants the archipelago would have migrate and closely consecutive record their modifications could preserved any one formation very many the marine inhabitants the archipelago now range thousands miles beyond its confines and analogy leads believe that would chiefly these far ranging species which would oftenest produce new varieties and the varieties would first generally local confined one place but possessed any decided advantage when further modified and improved they would slowly spread and supplant their parent forms when such varieties returned their ancient homes they would differ from their former state nearly uniform though perhaps extremely slight degree they would according the principles followed many palæontologists ranked new and distinct species then there some degree truth these remarks have right expect find our geological formations infinite number those fine transitional forms which theory assuredly have connected all the past and present species the same group into one long and branching chain life ought only look for few links some more closely some more distantly related each other and these links let them ever close found different stages the same formation would most palæontologists ranked distinct species but not pretend that should ever have suspected how poor record the mutations life the best preserved geological section presented had not the difficulty our not discovering innumerable transitional links between the species which appeared the commencement and close each formation pressed hardly theory the sudden appearance whole groups allied species the abrupt manner which whole groups species suddenly appear certain formations has been urged several palæontologists for instance agassiz pictet and none more forcibly than professor sedgwick fatal objection the belief the transmutation species numerous species belonging the same genera families have really started into life all once the fact would fatal the theory descent with slow modification through natural selection for the development group forms all which have descended from some one progenitor must have been extremely slow process and the progenitors must have lived long ages before their modified descendants but continually over rate the perfection the geological record and falsely infer because certain genera families have not been found beneath certain stage that they did not exist before that stage continually forget how large the world compared with the area over which our geological formations have been carefully examined forget that groups species may elsewhere have long existed and have slowly multiplied before they invaded the ancient archipelagoes europe and the united states not make due allowance for the enormous intervals time which have probably elapsed between our consecutive formations longer perhaps some cases than the time required for the accumulation each formation these intervals will have given time for the multiplication species from some one some few parent forms and the succeeding formation such species will appear suddenly created may here recall remark formerly made namely that might require long succession ages adapt organism some new and peculiar line life for instance fly through the air but that when this had been effected and few species had thus acquired great advantage over other organisms comparatively short time would necessary produce many divergent forms which would able spread rapidly and widely throughout the world will now give few examples illustrate these remarks and show how liable are error supposing that whole groups species have suddenly been produced may recall the well known fact that geological treatises published not many years ago the great class mammals was always spoken having abruptly come the commencement the tertiary series and now one the richest known accumulations fossil mammals belongs the middle the secondary series and one true mammal has been discovered the new red sandstone nearly the commencement this great series cuvier used urge that monkey occurred any tertiary stratum but now extinct species have been discovered india south america and europe even far back the eocene stage the most striking case however that the whale family these animals have huge bones are marine and range over the world the fact not single bone whale having been discovered any secondary formation seemed fully justify the belief that this great and distinct order had been suddenly produced the interval between the latest secondary and earliest tertiary formation but now may read the supplement lyell manual published 1858 clear evidence the existence whales the upper greensand some time before the close the secondary period may give another instance which from having passed under own eyes has much struck memoir fossil sessile cirripedes have stated that from the number existing and extinct tertiary species from the extraordinary abundance the individuals many species all over the world from the arctic regions the equator inhabiting various zones depths from the upper tidal limits fathoms from the perfect manner which specimens are preserved the oldest tertiary beds from the ease with which even fragment valve can recognised from all these circumstances inferred that had sessile cirripedes existed during the secondary periods they would certainly have been preserved and discovered and not one species had been discovered beds this age concluded that this great group had been suddenly developed the commencement the tertiary series this was sore trouble adding thought one more instance the abrupt appearance great group species but work had hardly been published when skilful palæontologist bosquet sent drawing perfect specimen unmistakeable sessile cirripede which had himself extracted from the chalk belgium and make the case striking possible this sessile cirripede was chthamalus very common large and ubiquitous genus which not one specimen has yet been found even any tertiary stratum hence now positively know that sessile cirripedes existed during the secondary period and these cirripedes might have been the progenitors our many tertiary and existing species the case most frequently insisted palæontologists the apparently sudden appearance whole group species that the teleostean fishes low down the chalk period this group includes the large majority existing species lately professor pictet has carried their existence one sub stage further back and some palæontologists believe that certain much older fishes which the affinities are yet imperfectly known are really teleostean assuming however that the whole them did appear agassiz believes the commencement the chalk formation the fact would certainly highly remarkable but cannot see that would insuperable difficulty theory unless could likewise shown that the species this group appeared suddenly and simultaneously throughout the world this same period almost superfluous remark that hardly any fossil fish are known from south the equator and running through pictet palæontology will seen that very few species are known from several formations europe some few families fish now have confined range the teleostean fish might formerly have had similarly confined range and after having been largely developed some one sea might have spread widely nor have any right suppose that the seas the world have always been freely open from south north they are present even this day the malay archipelago were converted into land the tropical parts the indian ocean would form large and perfectly enclosed basin which any great group marine animals might multiplied and here they would remain confined until some the species became adapted cooler climate and were enabled double the southern capes africa australia and thus reach other and distant seas from these and similar considerations but chiefly from our ignorance the geology other countries beyond the confines europe and the united states and from the revolution our palæontological ideas many points which the discoveries even the last dozen years have effected seems about rash dogmatize the succession organic beings throughout the world would for naturalist land for five minutes some one barren point australia and then discuss the number and range its productions the sudden appearance groups allied species the lowest known fossiliferous strata there another and allied difficulty which much graver allude the manner which numbers species the same group suddenly appear the lowest known fossiliferous rocks most the arguments which have convinced that all the existing species the same group have descended from one progenitor apply with nearly equal force the earliest known species for instance cannot doubt that all the silurian trilobites have descended from some one crustacean which must have lived long before the silurian age and which probably differed greatly from any known animal some the most ancient silurian animals the nautilus lingula etc not differ much from living species and cannot theory supposed that these old species were the progenitors all the species the orders which they belong for they not present characters any degree intermediate between them moreover they had been the progenitors these orders they would almost certainly have been long ago supplanted and exterminated their numerous and improved descendants consequently theory true indisputable that before the lowest silurian stratum was deposited long periods elapsed long probably far longer than the whole interval from the silurian age the present day and that during these vast yet quite unknown periods time the world swarmed with living creatures the question why not find records these vast primordial periods can give satisfactory answer several the most eminent geologists with sir murchison their head are convinced that see the organic remains the lowest silurian stratum the dawn life this planet other highly competent judges lyell and the late forbes dispute this conclusion should not forget that only small portion the world known with accuracy barrande has lately added another and lower stage the silurian system abounding with new and peculiar species traces life have been detected the longmynd beds beneath barrande called primordial zone the presence phosphatic nodules and bituminous matter some the lowest azoic rocks probably indicates the former existence life these periods but the difficulty understanding the absence vast piles fossiliferous strata which theory doubt were somewhere accumulated before the silurian epoch very great these most ancient beds had been wholly worn away denudation obliterated metamorphic action ought find only small remnants the formations next succeeding them age and these ought very generally metamorphosed condition but the descriptions which now possess the silurian deposits over immense territories russia and north america not support the view that the older formation the more has suffered the extremity denudation and metamorphism the case present must remain inexplicable and may truly urged valid argument against the views here entertained show that may hereafter receive some explanation will give the following hypothesis from the nature the organic remains which not appear have inhabited profound depths the several formations europe and the united states and from the amount sediment miles thickness which the formations are composed may infer that from first last large islands tracts land whence the sediment was derived occurred the neighbourhood the existing continents europe and north america but not know what was the state things the intervals between the successive formations whether europe and the united states during these intervals existed dry land submarine surface near land which sediment was not deposited again the bed open and unfathomable sea looking the existing oceans which are thrice extensive the land see them studded with many islands but not one oceanic island yet known afford even remnant any palæozoic secondary formation hence may perhaps infer that during the palæozoic and secondary periods neither continents nor continental islands existed where our oceans now extend for had they existed there palæozoic and secondary formations would all probability have been accumulated from sediment derived from their wear and tear and would have been least partially upheaved the oscillations level which may fairly conclude must have intervened during these enormously long periods then may infer anything from these facts may infer that where our oceans now extend oceans have extended from the remotest period which have any record and the other hand that where continents now exist large tracts land have existed subjected doubt great oscillations level since the earliest silurian period the coloured map appended volume coral reefs led conclude that the great oceans are still mainly areas subsidence the great archipelagoes still areas oscillations level and the continents areas elevation but have any right assume that things have thus remained from eternity our continents seem have been formed preponderance during many oscillations level the force elevation but may not the areas preponderant movement have changed the lapse ages period immeasurably antecedent the silurian epoch continents may have existed where oceans are now spread out and clear and open oceans may have existed where our continents now stand nor should justified assuming that for instance the bed the pacific ocean were now converted into continent should there find formations older than the silurian strata supposing such have been formerly deposited for might well happen that strata which had subsided some miles nearer the centre the earth and which had been pressed enormous weight superincumbent water might have undergone far more metamorphic action than strata which have always remained nearer the surface the immense areas some parts the world for instance south america bare metamorphic rocks which must have been heated under great pressure have always seemed require some special explanation and may perhaps believe that see these large areas the many formations long anterior the silurian epoch completely metamorphosed condition the several difficulties here discussed namely our not finding the successive formations infinitely numerous transitional links between the many species which now exist have existed the sudden manner which whole groups species appear our european formations the almost entire absence present known fossiliferous formations beneath the silurian strata are all undoubtedly the gravest nature see this the plainest manner the fact that all the most eminent palæontologists namely cuvier owen agassiz barrande falconer forbes etc and all our greatest geologists lyell murchison sedgwick etc have unanimously often vehemently maintained the immutability species but have reason believe that one great authority sir charles lyell from further reflexion entertains grave doubts this subject feel how rash differ from these great authorities whom with others owe all our knowledge those who think the natural geological record any degree perfect and who not attach much weight the facts and arguments other kinds given this volume will undoubtedly once reject theory for part following out lyell metaphor look the natural geological record history the world imperfectly kept and written changing dialect this history possess the last volume alone relating only two three countries this volume only here and there short chapter has been preserved and each page only here and there few lines each word the slowly changing language which the history supposed written being more less different the interrupted succession chapters may represent the apparently abruptly changed forms life entombed our consecutive but widely separated formations this view the difficulties above discussed are greatly diminished even disappear chapter the geological succession organic beings the slow and successive appearance new species their different rates change species once lost not reappear groups species follow the same general rules their appearance and disappearance single species extinction simultaneous changes the forms life throughout the world the affinities extinct species each other and living species the state development ancient forms the succession the same types within the same areas summary preceding and present chapters let now see whether the several facts and rules relating the geological succession organic beings better accord with the common view the immutability species with that their slow and gradual modification through descent and natural selection new species have appeared very slowly one after another both the land and the waters lyell has shown that hardly possible resist the evidence this head the case the several tertiary stages and every year tends fill the blanks between them and make the percentage system lost and new forms more gradual some the most recent beds though undoubtedly high antiquity measured years only one two species are lost forms and only one two are new forms having here appeared for the first time either locally far know the face the earth may trust the observations philippi sicily the successive changes the marine inhabitants that island have been many and most gradual the secondary formations are more broken but bronn has remarked neither the appearance nor disappearance their many now extinct species has been simultaneous each separate formation species different genera and classes have not changed the same rate the same degree the oldest tertiary beds few living shells may still found the midst multitude extinct forms falconer has given striking instance similar fact existing crocodile associated with many strange and lost mammals and reptiles the sub himalayan deposits the silurian lingula differs but little from the living species this genus whereas most the other silurian molluscs and all the crustaceans have changed greatly the productions the land seem change quicker rate than those the sea which striking instance has lately been observed switzerland there some reason believe that organisms considered high the scale nature change more quickly than those that are low though there are exceptions this rule the amount organic change pictet has remarked does not strictly correspond with the succession our geological formations that between each two consecutive formations the forms life have seldom changed exactly the same degree yet compare any but the most closely related formations all the species will found have undergone some change when species has once disappeared from the face the earth have reason believe that the same identical form never reappears the strongest apparent exception this latter rule that the called colonies barrande which intrude for period the midst older formation and then allow the pre existing fauna reappear but lyell explanation namely that case temporary migration from distinct geographical province seems satisfactory these several facts accord well with theory believe fixed law development causing all the inhabitants country change abruptly simultaneously equal degree the process modification must extremely slow the variability each species quite independent that all others whether such variability taken advantage natural selection and whether the variations accumulated greater lesser amount thus causing greater lesser amount modification the varying species depends many complex contingencies the variability being beneficial nature the power intercrossing the rate breeding the slowly changing physical conditions the country and more especially the nature the other inhabitants with which the varying species comes into competition hence means surprising that one species should retain the same identical form much longer than others changing that should change less see the same fact geographical distribution for instance the land shells and coleopterous insects madeira having come differ considerably from their nearest allies the continent europe whereas the marine shells and birds have remained unaltered can perhaps understand the apparently quicker rate change terrestrial and more highly organised productions compared with marine and lower productions the more complex relations the higher beings their organic and inorganic conditions life explained former chapter when many the inhabitants country have become modified and improved can understand the principle competition and that the many all important relations organism organism that any form which does not become some degree modified and improved will liable exterminated hence can see why all the species the same region last look wide enough intervals time become modified for those which not change will become extinct members the same class the average amount change during long and equal periods time may perhaps nearly the same but the accumulation long enduring fossiliferous formations depends great masses sediment having been deposited areas whilst subsiding our formations have been almost necessarily accumulated wide and irregularly intermittent intervals consequently the amount organic change exhibited the fossils embedded consecutive formations not equal each formation this view does not mark new and complete act creation but only occasional scene taken almost hazard slowly changing drama can clearly understand why species when once lost should never reappear even the very same conditions life organic and inorganic should recur for though the offspring one species might adapted and doubt this has occurred innumerable instances fill the exact place another species the economy nature and thus supplant yet the two forms the old and the new would not identically the same for both would almost certainly inherit different characters from their distinct progenitors for instance just possible our fantail pigeons were all destroyed that fanciers striving during long ages for the same object might make new breed hardly distinguishable from our present fantail but the parent rock pigeon were also destroyed and nature have every reason believe that the parent form will generally supplanted and exterminated its improved offspring quite incredible that fantail identical with the existing breed could raised from any other species pigeon even from the other well established races the domestic pigeon for the newly formed fantail would almost sure inherit from its new progenitor some slight characteristic differences groups species that genera and families follow the same general rules their appearance and disappearance single species changing more less quickly and greater lesser degree group does not reappear after has once disappeared its existence long lasts continuous aware that there are some apparent exceptions this rule but the exceptions are surprisingly few few that forbes pictet and woodward though all strongly opposed such views maintain admit its truth and the rule strictly accords with theory for all the species the same group have descended from some one species clear that long any species the group have appeared the long succession ages long must its members have continuously existed order have generated either new and modified the same old and unmodified forms species the genus lingula for instance must have continuously existed unbroken succession generations from the lowest silurian stratum the present day have seen the last chapter that the species group sometimes falsely appear have come abruptly and have attempted give explanation this fact which true would have been fatal views but such cases are certainly exceptional the general rule being gradual increase number till the group reaches its maximum and then sooner later gradually decreases the number the species genus the number the genera family represented vertical line varying thickness crossing the successive geological formations which the species are found the line will sometimes falsely appear begin its lower end not sharp point but abruptly then gradually thickens upwards sometimes keeping for space equal thickness and ultimately thins out the upper beds marking the decrease and final extinction the species this gradual increase number the species group strictly conformable with theory the species the same genus and the genera the same family can increase only slowly and progressively for the process modification and the production number allied forms must slow and gradual one species giving rise first two three varieties these being slowly converted into species which their turn produce equally slow steps other species and like the branching great tree from single stem till the group becomes large extinction have yet spoken only incidentally the disappearance species and groups species the theory natural selection the extinction old forms and the production new and improved forms are intimately connected together the old notion all the inhabitants the earth having been swept away successive periods catastrophes very generally given even those geologists elie beaumont murchison barrande etc whose general views would naturally lead them this conclusion the contrary have every reason believe from the study the tertiary formations that species and groups species gradually disappear one after another first from one spot then from another and finally from the world both single species and whole groups species last for very unequal periods some groups have seen having endured from the earliest known dawn life the present day some having disappeared before the close the palæozoic period fixed law seems determine the length time during which any single species any single genus endures there reason believe that the complete extinction the species group generally slower process than their production the appearance and disappearance group species represented before vertical line varying thickness the line found taper more gradually its upper end which marks the progress extermination than its lower end which marks the first appearance and increase numbers the species some cases however the extermination whole groups beings ammonites towards the close the secondary period has been wonderfully sudden the whole subject the extinction species has been involved the most gratuitous mystery some authors have even supposed that the individual has definite length life have species definite duration one think can have marvelled more the extinction species than have done when found plata the tooth horse embedded with the remains mastodon megatherium toxodon and other extinct monsters which all existed with still living shells very late geological period was filled with astonishment for seeing that the horse since its introduction the spaniards into south america has run wild over the whole country and has increased numbers unparalleled rate asked myself what could recently have exterminated the former horse under conditions life apparently favourable but how utterly groundless was astonishment professor owen soon perceived that the tooth though like that the existing horse belonged extinct species had this horse been still living but some degree rare naturalist would have felt the least surprise its rarity for rarity the attribute vast number species all classes all countries ask ourselves why this that species rare answer that something unfavourable its conditions life but what that something can hardly ever tell the supposition the fossil horse still existing rare species might have felt certain from the analogy all other mammals even the slow breeding elephant and from the history the naturalisation the domestic horse south america that under more favourable conditions would very few years have stocked the whole continent but could not have told what the unfavourable conditions were which checked its increase whether some one several contingencies and what period the horse life and what degree they severally acted the conditions had gone however slowly becoming less and less favourable assuredly should not have perceived the fact yet the fossil horse would certainly have become rarer and rarer and finally extinct its place being seized some more successful competitor most difficult always remember that the increase every living being constantly being checked unperceived injurious agencies and that these same unperceived agencies are amply sufficient cause rarity and finally extinction see many cases the more recent tertiary formations that rarity precedes extinction and know that this has been the progress events with those animals which have been exterminated either locally wholly through man agency may repeat what published 1845 namely that admit that species generally become rare before they become extinct feel surprise the rarity species and yet marvel greatly when ceases exist much the same admit that sickness the individual the forerunner death feel surprise sickness but when the sick man dies wonder and suspect that died some unknown deed violence the theory natural selection grounded the belief that each new variety and ultimately each new species produced and maintained having some advantage over those with which comes into competition and the consequent extinction less favoured forms almost inevitably follows the same with our domestic productions when new and slightly improved variety has been raised first supplants the less improved varieties the same neighbourhood when much improved transported far and near like our short horn cattle and takes the place other breeds other countries thus the appearance new forms and the disappearance old forms both natural and artificial are bound together certain flourishing groups the number new specific forms which have been produced within given time probably greater than that the old forms which have been exterminated but know that the number species has not gone indefinitely increasing least during the later geological periods that looking later times may believe that the production new forms has caused the extinction about the same number old forms the competition will generally most severe formerly explained and illustrated examples between the forms which are most like each other all respects hence the improved and modified descendants species will generally cause the extermination the parent species and many new forms have been developed from any one species the nearest allies that species the species the same genus will the most liable extermination thus believe number new species descended from one species that new genus comes supplant old genus belonging the same family but must often have happened that new species belonging some one group will have seized the place occupied species belonging distinct group and thus caused its extermination and many allied forms developed from the successful intruder many will have yield their places and will generally allied forms which will suffer from some inherited inferiority common but whether species belonging the same distinct class which yield their places other species which have been modified and improved few the sufferers may often long preserved from being fitted some peculiar line life from inhabiting some distant and isolated station where they have escaped severe competition for instance single species trigonia great genus shells the secondary formations survives the australian seas and few members the great and almost extinct group ganoid fishes still inhabit our fresh waters therefore the utter extinction group generally have seen slower process than its production with respect the apparently sudden extermination whole families orders trilobites the close the palæozoic period and ammonites the close the secondary period must remember what has been already said the probable wide intervals time between our consecutive formations and these intervals there may have been much slow extermination moreover when sudden immigration unusually rapid development many species new group have taken possession new area they will have exterminated correspondingly rapid manner many the old inhabitants and the forms which thus yield their places will commonly allied for they will partake some inferiority common thus seems the manner which single species and whole groups species become extinct accords well with the theory natural selection need not marvel extinction must marvel let our presumption imagining for moment that understand the many complex contingencies which the existence each species depends forget for instant that each species tends increase inordinately and that some check always action yet seldom perceived the whole economy nature will utterly obscured whenever can precisely say why this species more abundant individuals than that why this species and not another can naturalised given country then and not till then may justly feel surprise why cannot account for the extinction this particular species group species the forms life changing almost simultaneously throughout the world scarcely any palæontological discovery more striking than the fact that the forms life change almost simultaneously throughout the world thus our european chalk formation can recognised many distant parts the world under the most different climates where not fragment the mineral chalk itself can found namely north america equatorial south america tierra del fuego the cape good hope and the peninsula india for these distant points the organic remains certain beds present unmistakeable degree resemblance those the chalk not that the same species are met with for some cases not one species identically the same but they belong the same families genera and sections genera and sometimes are similarly characterised such trifling points mere superficial sculpture moreover other forms which are not found the chalk europe but which occur the formations either above below are similarly absent these distant points the world the several successive palæozoic formations russia western europe and north america similar parallelism the forms life has been observed several authors according lyell with the several european and north american tertiary deposits even the few fossil species which are common the old and new worlds kept wholly out view the general parallelism the successive forms life the stages the widely separated palæozoic and tertiary periods would still manifest and the several formations could easily correlated these observations however relate the marine inhabitants distant parts the world have not sufficient data judge whether the productions the land and fresh water change distant points the same parallel manner may doubt whether they have thus changed the megatherium mylodon macrauchenia and toxodon had been brought europe from plata without any information regard their geological position one would have suspected that they had coexisted with still living sea shells but these anomalous monsters coexisted with the mastodon and horse might least have been inferred that they had lived during one the latter tertiary stages when the marine forms life are spoken having changed simultaneously throughout the world must not supposed that this expression relates the same thousandth hundred thousandth year even that has very strict geological sense for all the marine animals which live the present day europe and all those that lived europe during the pleistocene period enormously remote period measured years including the whole glacial epoch were compared with those now living south america australia the most skilful naturalist would hardly able say whether the existing the pleistocene inhabitants europe resembled most closely those the southern hemisphere again several highly competent observers believe that the existing productions the united states are more closely related those which lived europe during certain later tertiary stages than those which now live here and this evident that fossiliferous beds deposited the present day the shores north america would hereafter liable classed with somewhat older european beds nevertheless looking remotely future epoch there can think little doubt that all the more modern marine formations namely the upper pliocene the pleistocene and strictly modern beds europe north and south america and australia from containing fossil remains some degree allied and from not including those forms which are only found the older underlying deposits would correctly ranked simultaneous geological sense the fact the forms life changing simultaneously the above large sense distant parts the world has greatly struck those admirable observers verneuil and archiac after referring the parallelism the palæozoic forms life various parts europe they add struck this strange sequence turn our attention north america and there discover series analogous phenomena will appear certain that all these modifications species their extinction and the introduction new ones cannot owing mere changes marine currents other causes more less local and temporary but depend general laws which govern the whole animal kingdom barrande has made forcible remarks precisely the same effect indeed quite futile look changes currents climate other physical conditions the cause these great mutations the forms life throughout the world under the most different climates must barrande has remarked look some special law shall see this more clearly when treat the present distribution organic beings and find how slight the relation between the physical conditions various countries and the nature their inhabitants this great fact the parallel succession the forms life throughout the world explicable the theory natural selection new species are formed new varieties arising which have some advantage over older forms and those forms which are already dominant have some advantage over the other forms their own country would naturally oftenest give rise new varieties incipient species for these latter must victorious still higher degree order preserved and survive have distinct evidence this head the plants which are dominant that which are commonest their own homes and are most widely diffused having produced the greatest number new varieties also natural that the dominant varying and far spreading species which already have invaded certain extent the territories other species should those which would have the best chance spreading still further and giving rise new countries new varieties and species the process diffusion may often very slow being dependent climatal and geographical changes strange accidents but the long run the dominant forms will generally succeed spreading the diffusion would probable slower with the terrestrial inhabitants distinct continents than with the marine inhabitants the continuous sea might therefore expect find apparently find less strict degree parallel succession the productions the land than the sea dominant species spreading from any region might encounter still more dominant species and then their triumphant course even their existence would cease know not all precisely what are all the conditions most favourable for the multiplication new and dominant species but can think clearly see that number individuals from giving better chance the appearance favourable variations and that severe competition with many already existing forms would highly favourable would the power spreading into new territories certain amount isolation recurring long intervals time would probably also favourable before explained one quarter the world may have been most favourable for the production new and dominant species the land and another for those the waters the sea two great regions had been for long period favourably circumstanced equal degree whenever their inhabitants met the battle would prolonged and severe and some from one birthplace and some from the other might victorious but the course time the forms dominant the highest degree wherever produced would tend everywhere prevail they prevailed they would cause the extinction other and inferior forms and these inferior forms would allied groups inheritance whole groups would tend slowly disappear though here and there single member might long enabled survive thus seems the parallel and taken large sense simultaneous succession the same forms life throughout the world accords well with the principle new species having been formed dominant species spreading widely and varying the new species thus produced being themselves dominant owing inheritance and having already had some advantage over their parents over other species these again spreading varying and producing new species the forms which are beaten and which yield their places the new and victorious forms will generally allied groups from inheriting some inferiority common and therefore new and improved groups spread throughout the world old groups will disappear from the world and the succession forms both ways will everywhere tend correspond there one other remark connected with this subject worth making have given reasons for believing that all our greater fossiliferous formations were deposited during periods subsidence and that blank intervals vast duration occurred during the periods when the bed the sea was either stationary rising and likewise when sediment was not thrown down quickly enough embed and preserve organic remains during these long and blank intervals suppose that the inhabitants each region underwent considerable amount modification and extinction and that there was much migration from other parts the world have reason believe that large areas are affected the same movement probable that strictly contemporaneous formations have often been accumulated over very wide spaces the same quarter the world but are far from having any right conclude that this has invariably been the case and that large areas have invariably been affected the same movements when two formations have been deposited two regions during nearly but not exactly the same period should find both from the causes explained the foregoing paragraphs the same general succession the forms life but the species would not exactly correspond for there will have been little more time the one region than the other for modification extinction and immigration suspect that cases this nature have occurred europe prestwich his admirable memoirs the eocene deposits england and france able draw close general parallelism between the successive stages the two countries but when compares certain stages england with those france although finds both curious accordance the numbers the species belonging the same genera yet the species themselves differ manner very difficult account for considering the proximity the two areas unless indeed assumed that isthmus separated two seas inhabited distinct but contemporaneous faunas lyell has made similar observations some the later tertiary formations barrande also shows that there striking general parallelism the successive silurian deposits bohemia and scandinavia nevertheless finds surprising amount difference the species the several formations these regions have not been deposited during the same exact periods formation one region often corresponding with blank interval the other and both regions the species have gone slowly changing during the accumulation the several formations and during the long intervals time between them this case the several formations the two regions could arranged the same order accordance with the general succession the form life and the order would falsely appear strictly parallel nevertheless the species would not all the same the apparently corresponding stages the two regions the affinities extinct species each other and living forms let now look the mutual affinities extinct and living species they all fall into one grand natural system and this fact once explained the principle descent the more ancient any form the more general rule differs from living forms but buckland long ago remarked all fossils can classed either still existing groups between them that the extinct forms life help fill the wide intervals between existing genera families and orders cannot disputed for confine our attention either the living the extinct alone the series far less perfect than combine both into one general system with respect the vertebrata whole pages could filled with striking illustrations from our great palæontologist owen showing how extinct animals fall between existing groups cuvier ranked the ruminants and pachyderms the two most distinct orders mammals but owen has discovered many fossil links that has had alter the whole classification these two orders and has placed certain pachyderms the same sub order with ruminants for example dissolves fine gradations the apparently wide difference between the pig and the camel regard the invertebrata barrande and higher authority could not named asserts that every day taught that palæozoic animals though belonging the same orders families genera with those living the present day were not this early epoch limited such distinct groups they now are some writers have objected any extinct species group species being considered intermediate between living species groups this term meant that extinct form directly intermediate all its characters between two living forms the objection probably valid but apprehend that perfectly natural classification many fossil species would have stand between living species and some extinct genera between living genera even between genera belonging distinct families the most common case especially with respect very distinct groups such fish and reptiles seems that supposing them distinguished the present day from each other dozen characters the ancient members the same two groups would distinguished somewhat lesser number characters that the two groups though formerly quite distinct that period made some small approach each other common belief that the more ancient form much the more tends connect some its characters groups now widely separated from each other this remark doubt must restricted those groups which have undergone much change the course geological ages and would difficult prove the truth the proposition for every now and then even living animal the lepidosiren discovered having affinities directed towards very distinct groups yet compare the older reptiles and batrachians the older fish the older cephalopods and the eocene mammals with the more recent members the same classes must admit that there some truth the remark let see how far these several facts and inferences accord with the theory descent with modification the subject somewhat complex must request the reader turn the diagram the fourth chapter may suppose that the numbered letters represent genera and the dotted lines diverging from them the species each genus the diagram much too simple too few genera and too few species being given but this unimportant for the horizontal lines may represent successive geological formations and all the forms beneath the uppermost line may considered extinct the three existing genera will form small family and closely allied family sub family and third family these three families together with the many extinct genera the several lines descent diverging from the parent form will form order for all will have inherited something common from their ancient and common progenitor the principle the continued tendency divergence character which was formerly illustrated this diagram the more recent any form the more will generally differ from its ancient progenitor hence can understand the rule that the most ancient fossils differ most from existing forms must not however assume that divergence character necessary contingency depends solely the descendants from species being thus enabled seize many and different places the economy nature therefore quite possible have seen the case some silurian forms that species might being slightly modified relation its slightly altered conditions life and yet retain throughout vast period the same general characteristics this represented the diagram the letter f14 all the many forms extinct and recent descended from make before remarked one order and this order from the continued effects extinction and divergence character has become divided into several sub families and families some which are supposed have perished different periods and some have endured the present day looking the diagram can see that many the extinct forms supposed embedded the successive formations were discovered several points low down the series the three existing families the uppermost line would rendered less distinct from each other for instance the genera were disinterred these three families would closely linked together that they probably would have united into one great family nearly the same manner has occurred with ruminants and pachyderms yet who objected call the extinct genera which thus linked the living genera three families together intermediate character would justified they are intermediate not directly but only long and circuitous course through many widely different forms many extinct forms were discovered above one the middle horizontal lines geological formations for instance above number but none from beneath this line then only the two families the left hand namely etc and etc would have united into one family and the two other families namely now including five genera and would yet remain distinct these two families however would less distinct from each other than they were before the discovery the fossils for instance suppose the existing genera the two families differ from each other dozen characters this case the genera the early period marked would differ lesser number characters for this early stage descent they have not diverged character from the common progenitor the order nearly much they subsequently diverged thus comes that ancient and extinct genera are often some slight degree intermediate character between their modified descendants between their collateral relations nature the case will far more complicated than represented the diagram for the groups will have been more numerous they will have endured for extremely unequal lengths time and will have been modified various degrees possess only the last volume the geological record and that very broken condition have right expect except very rare cases fill wide intervals the natural system and thus unite distinct families orders all that have right expect that those groups which have within known geological periods undergone much modification should the older formations make some slight approach each other that the older members should differ less from each other some their characters than the existing members the same groups and this the concurrent evidence our best palæontologists seems frequently the case thus the theory descent with modification the main facts with respect the mutual affinities the extinct forms life each other and living forms seem explained satisfactory manner and they are wholly inexplicable any other view this same theory evident that the fauna any great period the earth history will intermediate general character between that which preceded and that which succeeded thus the species which lived the sixth great stage descent the diagram are the modified offspring those which lived the fifth stage and are the parents those which became still more modified the seventh stage hence they could hardly fail nearly intermediate character between the forms life above and below must however allow for the entire extinction some preceding forms and for the coming quite new forms immigration and for large amount modification during the long and blank intervals between the successive formations subject these allowances the fauna each geological period undoubtedly intermediate character between the preceding and succeeding faunas need give only one instance namely the manner which the fossils the devonian system when this system was first discovered were once recognised palæontologists intermediate character between those the overlying carboniferous and underlying silurian system but each fauna not necessarily exactly intermediate unequal intervals time have elapsed between consecutive formations real objection the truth the statement that the fauna each period whole nearly intermediate character between the preceding and succeeding faunas that certain genera offer exceptions the rule for instance mastodons and elephants when arranged falconer two series first according their mutual affinities and then according their periods existence not accord arrangement the species extreme character are not the oldest the most recent nor are those which are intermediate character intermediate age but supposing for instant this and other such cases that the record the first appearance and disappearance the species was perfect have reason believe that forms successively produced necessarily endure for corresponding lengths time very ancient form might occasionally last much longer than form elsewhere subsequently produced especially the case terrestrial productions inhabiting separated districts compare small things with great the principal living and extinct races the domestic pigeon were arranged well they could serial affinity this arrangement would not closely accord with the order time their production and still less with the order their disappearance for the parent rock pigeon now lives and many varieties between the rock pigeon and the carrier have become extinct and carriers which are extreme the important character length beak originated earlier than short beaked tumblers which are the opposite end the series this same respect closely connected with the statement that the organic remains from intermediate formation are some degree intermediate character the fact insisted all palæontologists that fossils from two consecutive formations are far more closely related each other than are the fossils from two remote formations pictet gives well known instance the general resemblance the organic remains from the several stages the chalk formation though the species are distinct each stage this fact alone from its generality seems have shaken professor pictet his firm belief the immutability species who acquainted with the distribution existing species over the globe will not attempt account for the close resemblance the distinct species closely consecutive formations the physical conditions the ancient areas having remained nearly the same let remembered that the forms life least those inhabiting the sea have changed almost simultaneously throughout the world and therefore under the most different climates and conditions consider the prodigious vicissitudes climate during the pleistocene period which includes the whole glacial period and note how little the specific forms the inhabitants the sea have been affected the theory descent the full meaning the fact fossil remains from closely consecutive formations though ranked distinct species being closely related obvious the accumulation each formation has often been interrupted and long blank intervals have intervened between successive formations ought not expect find attempted show the last chapter any one two formations all the intermediate varieties between the species which appeared the commencement and close these periods but ought find after intervals very long measured years but only moderately long measured geologically closely allied forms they have been called some authors representative species and these assuredly find find short such evidence the slow and scarcely sensible mutation specific forms have just right expect find the state development ancient forms there has been much discussion whether recent forms are more highly developed than ancient will not here enter this subject for naturalists have not yet defined each other satisfaction what meant high and low forms but one particular sense the more recent forms must theory higher than the more ancient for each new species formed having had some advantage the struggle for life over other and preceding forms under nearly similar climate the eocene inhabitants one quarter the world were put into competition with the existing inhabitants the same some other quarter the eocene fauna flora would certainly beaten and exterminated would secondary fauna eocene and palæozoic fauna secondary fauna not doubt that this process improvement has affected marked and sensible manner the organisation the more recent and victorious forms life comparison with the ancient and beaten forms but can see way testing this sort progress crustaceans for instance not the highest their own class may have beaten the highest molluscs from the extraordinary manner which european productions have recently spread over new zealand and have seized places which must have been previously occupied may believe all the animals and plants great britain were set free new zealand that the course time multitude british forms would become thoroughly naturalized there and would exterminate many the natives the other hand from what see now occurring new zealand and from hardly single inhabitant the southern hemisphere having become wild any part europe may doubt all the productions new zealand were set free great britain whether any considerable number would enabled seize places now occupied our native plants and animals under this point view the productions great britain may said higher than those new zealand yet the most skilful naturalist from examination the species the two countries could not have foreseen this result agassiz insists that ancient animals resemble certain extent the embryos recent animals the same classes that the geological succession extinct forms some degree parallel the embryological development recent forms must follow pictet and huxley thinking that the truth this doctrine very far from proved yet fully expect see hereafter confirmed least regard subordinate groups which have branched off from each other within comparatively recent times for this doctrine agassiz accords well with the theory natural selection future chapter shall attempt show that the adult differs from its embryo owing variations supervening not early age and being inherited corresponding age this process whilst leaves the embryo almost unaltered continually adds the course successive generations more and more difference the adult thus the embryo comes left sort picture preserved nature the ancient and less modified condition each animal this view may true and yet may never capable full proof seeing for instance that the oldest known mammals reptiles and fish strictly belong their own proper classes though some these old forms are slight degree less distinct from each other than are the typical members the same groups the present day would vain look for animals having the common embryological character the vertebrata until beds far beneath the lowest silurian strata are discovered discovery which the chance very small the succession the same types within the same areas during the later tertiary periods clift many years ago showed that the fossil mammals from the australian caves were closely allied the living marsupials that continent south america similar relationship manifest even uneducated eye the gigantic pieces armour like those the armadillo found several parts plata and professor owen has shown the most striking manner that most the fossil mammals buried there such numbers are related south american types this relationship even more clearly seen the wonderful collection fossil bones made lund and clausen the caves brazil was much impressed with these facts that strongly insisted 1839 and 1845 this law the succession types this wonderful relationship the same continent between the dead and the living professor owen has subsequently extended the same generalisation the mammals the old world see the same law this author restorations the extinct and gigantic birds new zealand see also the birds the caves brazil woodward has shown that the same law holds good with sea shells but from the wide distribution most genera molluscs not well displayed them other cases could added the relation between the extinct and living land shells madeira and between the extinct and living brackish water shells the aralo caspian sea now what does this remarkable law the succession the same types within the same areas mean would bold man who after comparing the present climate australia and parts south america under the same latitude would attempt account the one hand dissimilar physical conditions for the dissimilarity the inhabitants these two continents and the other hand similarity conditions for the uniformity the same types each during the later tertiary periods nor can pretended that immutable law that marsupials should have been chiefly solely produced australia that edentata and other american types should have been solely produced south america for know that europe ancient times was peopled numerous marsupials and have shown the publications above alluded that america the law distribution terrestrial mammals was formerly different from what now north america formerly partook strongly the present character the southern half the continent and the southern half was formerly more closely allied than present the northern half similar manner know from falconer and cautley discoveries that northern india was formerly more closely related its mammals africa than the present time analogous facts could given relation the distribution marine animals the theory descent with modification the great law the long enduring but not immutable succession the same types within the same areas once explained for the inhabitants each quarter the world will obviously tend leave that quarter during the next succeeding period time closely allied though some degree modified descendants the inhabitants one continent formerly differed greatly from those another continent will their modified descendants still differ nearly the same manner and degree but after very long intervals time and after great geographical changes permitting much inter migration the feebler will yield the more dominant forms and there will nothing immutable the laws past and present distribution may asked ridicule whether suppose that the megatherium and other allied huge monsters have left behind them south america the sloth armadillo and anteater their degenerate descendants this cannot for instant admitted these huge animals have become wholly extinct and have left progeny but the caves brazil there are many extinct species which are closely allied size and other characters the species still living south america and some these fossils may the actual progenitors living species must not forgotten that theory all the species the same genus have descended from some one species that six genera each having eight species found one geological formation and the next succeeding formation there six other allied representative genera with the same number species then may conclude that only one species each the six older genera has left modified descendants constituting the six new genera the other seven species the old genera have all died out and have left progeny which would probably far commoner case two three species two three alone the six older genera will have been the parents the six new genera the other old species and the other whole genera having become utterly extinct failing orders with the genera and species decreasing numbers apparently the case the edentata south america still fewer genera and species will have left modified blood descendants summary the preceding and present chapters have attempted show that the geological record extremely imperfect that only small portion the globe has been geologically explored with care that only certain classes organic beings have been largely preserved fossil state that the number both specimens and species preserved our museums absolutely nothing compared with the incalculable number generations which must have passed away even during single formation that owing subsidence being necessary for the accumulation fossiliferous deposits thick enough resist future degradation enormous intervals time have elapsed between the successive formations that there has probably been more extinction during the periods subsidence and more variation during the periods elevation and during the latter the record will have been least perfectly kept that each single formation has not been continuously deposited that the duration each formation perhaps short compared with the average duration specific forms that migration has played important part the first appearance new forms any one area and formation that widely ranging species are those which have varied most and have oftenest given rise new species and that varieties have first often been local all these causes taken conjointly must have tended make the geological record extremely imperfect and will large extent explain why not find interminable varieties connecting together all the extinct and existing forms life the finest graduated steps who rejects these views the nature the geological record will rightly reject whole theory for may ask vain where are the numberless transitional links which must formerly have connected the closely allied representative species found the several stages the same great formation may disbelieve the enormous intervals time which have elapsed between our consecutive formations may overlook how important part migration must have played when the formations any one great region alone that europe are considered may urge the apparent but often falsely apparent sudden coming whole groups species may ask where are the remains those infinitely numerous organisms which must have existed long before the first bed the silurian system was deposited can answer this latter question only hypothetically saying that far can see where our oceans now extend they have for enormous period extended and where our oscillating continents now stand they have stood ever since the silurian epoch but that long before that period the world may have presented wholly different aspect and that the older continents formed formations older than any known may now all metamorphosed condition may lie buried under the ocean passing from these difficulties all the other great leading facts palæontology seem simply follow the theory descent with modification through natural selection can thus understand how that new species come slowly and successively how species different classes not necessarily change together the same rate the same degree yet the long run that all undergo modification some extent the extinction old forms the almost inevitable consequence the production new forms can understand why when species has once disappeared never reappears groups species increase numbers slowly and endure for unequal periods time for the process modification necessarily slow and depends many complex contingencies the dominant species the larger dominant groups tend leave many modified descendants and thus new sub groups and groups are formed these are formed the species the less vigorous groups from their inferiority inherited from common progenitor tend become extinct together and leave modified offspring the face the earth but the utter extinction whole group species may often very slow process from the survival few descendants lingering protected and isolated situations when group has once wholly disappeared does not reappear for the link generation has been broken can understand how the spreading the dominant forms life which are those that oftenest vary will the long run tend people the world with allied but modified descendants and these will generally succeed taking the places those groups species which are their inferiors the struggle for existence hence after long intervals time the productions the world will appear have changed simultaneously can understand how that all the forms life ancient and recent make together one grand system for all are connected generation can understand from the continued tendency divergence character why the more ancient form the more generally differs from those now living why ancient and extinct forms often tend fill gaps between existing forms sometimes blending two groups previously classed distinct into one but more commonly only bringing them little closer together the more ancient form the more often apparently displays characters some degree intermediate between groups now distinct for the more ancient form the more nearly will related and consequently resemble the common progenitor groups since become widely divergent extinct forms are seldom directly intermediate between existing forms but are intermediate only long and circuitous course through many extinct and very different forms can clearly see why the organic remains closely consecutive formations are more closely allied each other than are those remote formations for the forms are more closely linked together generation can clearly see why the remains intermediate formation are intermediate character the inhabitants each successive period the world history have beaten their predecessors the race for life and are far higher the scale nature and this may account for that vague yet ill defined sentiment felt many palæontologists that organisation the whole has progressed should hereafter proved that ancient animals resemble certain extent the embryos more recent animals the same class the fact will intelligible the succession the same types structure within the same areas during the later geological periods ceases mysterious and simply explained inheritance then the geological record imperfect believe and may least asserted that the record cannot proved much more perfect the main objections the theory natural selection are greatly diminished disappear the other hand all the chief laws palæontology plainly proclaim seems that species have been produced ordinary generation old forms having been supplanted new and improved forms life produced the laws variation still acting round and preserved natural selection chapter geographical distribution present distribution cannot accounted for differences physical conditions importance barriers affinity the productions the same continent centres creation means dispersal changes climate and the level the land and occasional means dispersal during the glacial period extensive with the world considering the distribution organic beings over the face the globe the first great fact which strikes that neither the similarity nor the dissimilarity the inhabitants various regions can accounted for their climatal and other physical conditions late almost every author who has studied the subject has come this conclusion the case america alone would almost suffice prove its truth for exclude the northern parts where the circumpolar land almost continuous all authors agree that one the most fundamental divisions geographical distribution that between the new and old worlds yet travel over the vast american continent from the central parts the united states its extreme southern point meet with the most diversified conditions the most humid districts arid deserts lofty mountains grassy plains forests marshes lakes and great rivers under almost every temperature there hardly climate condition the old world which cannot paralleled the new least closely the same species generally require for most rare case find group organisms confined any small spot having conditions peculiar only slight degree for instance small areas the old world could pointed out hotter than any the new world yet these are not inhabited peculiar fauna flora notwithstanding this parallelism the conditions the old and new worlds how widely different are their living productions the southern hemisphere compare large tracts land australia south africa and western south america between latitudes and shall find parts extremely similar all their conditions yet would not possible point out three faunas and floras more utterly dissimilar again may compare the productions south america south lat with those north which consequently inhabit considerably different climate and they will found incomparably more closely related each other than they are the productions australia africa under nearly the same climate analogous facts could given with respect the inhabitants the sea second great fact which strikes our general review that barriers any kind obstacles free migration are related close and important manner the differences between the productions various regions see this the great difference nearly all the terrestrial productions the new and old worlds excepting the northern parts where the land almost joins and where under slightly different climate there might have been free migration for the northern temperate forms there now for the strictly arctic productions see the same fact the great difference between the inhabitants australia africa and south america under the same latitude for these countries are almost much isolated from each other possible each continent also see the same fact for the opposite sides lofty and continuous mountain ranges and great deserts and sometimes even large rivers find different productions though mountain chains deserts etc are not impassable likely have endured long the oceans separating continents the differences are very inferior degree those characteristic distinct continents turning the sea find the same law two marine faunas are more distinct with hardly fish shell crab common than those the eastern and western shores south and central america yet these great faunas are separated only the narrow but impassable isthmus panama westward the shores america wide space open ocean extends with not island halting place for emigrants here have barrier another kind and soon this passed meet the eastern islands the pacific with another and totally distinct fauna that here three marine faunas range far northward and southward parallel lines not far from each other under corresponding climates but from being separated from each other impassable barriers either land open sea they are wholly distinct the other hand proceeding still further westward from the eastern islands the tropical parts the pacific encounter impassable barriers and have innumerable islands halting places until after travelling over hemisphere come the shores africa and over this vast space meet with well defined and distinct marine faunas although hardly one shell crab fish common the above named three approximate faunas eastern and western america and the eastern pacific islands yet many fish range from the pacific into the indian ocean and many shells are common the eastern islands the pacific and the eastern shores africa almost exactly opposite meridians longitude third great fact partly included the foregoing statements the affinity the productions the same continent sea though the species themselves are distinct different points and stations law the widest generality and every continent offers innumerable instances nevertheless the naturalist travelling for instance from north south never fails struck the manner which successive groups beings specifically distinct yet clearly related replace each other hears from closely allied yet distinct kinds birds notes nearly similar and sees their nests similarly constructed but not quite alike with eggs coloured nearly the same manner the plains near the straits magellan are inhabited one species rhea american ostrich and northward the plains plata another species the same genus and not true ostrich emeu like those found africa and australia under the same latitude these same plains plata see the agouti and bizcacha animals having nearly the same habits our hares and rabbits and belonging the same order rodents but they plainly display american type structure ascend the lofty peaks the cordillera and find alpine species bizcacha look the waters and not find the beaver musk rat but the coypu and capybara rodents the american type innumerable other instances could given look the islands off the american shore however much they may differ geological structure the inhabitants though they may all peculiar species are essentially american may look back past ages shown the last chapter and find american types then prevalent the american continent and the american seas see these facts some deep organic bond prevailing throughout space and time over the same areas land and water and independent their physical conditions the naturalist must feel little curiosity who not led inquire what this bond this bond theory simply inheritance that cause which alone far positively know produces organisms quite like see the case varieties nearly like each other the dissimilarity the inhabitants different regions may attributed modification through natural selection and quite subordinate degree the direct influence different physical conditions the degree dissimilarity will depend the migration the more dominant forms life from one region into another having been effected with more less ease periods more less remote the nature and number the former immigrants and their action and reaction their mutual struggles for life the relation organism organism being have already often remarked the most important all relations thus the high importance barriers comes into play checking migration does time for the slow process modification through natural selection widely ranging species abounding individuals which have already triumphed over many competitors their own widely extended homes will have the best chance seizing new places when they spread into new countries their new homes they will exposed new conditions and will frequently undergo further modification and improvement and thus they will become still further victorious and will produce groups modified descendants this principle inheritance with modification can understand how that sections genera whole genera and even families are confined the same areas commonly and notoriously the case believe was remarked the last chapter law necessary development the variability each species independent property and will taken advantage natural selection only far profits the individual its complex struggle for life the degree modification different species will uniform quantity for instance number species which stand direct competition with each other migrate body into new and afterwards isolated country they will little liable modification for neither migration nor isolation themselves can anything these principles come into play only bringing organisms into new relations with each other and lesser degree with the surrounding physical conditions have seen the last chapter that some forms have retained nearly the same character from enormously remote geological period certain species have migrated over vast spaces and have not become greatly modified these views obvious that the several species the same genus though inhabiting the most distant quarters the world must originally have proceeded from the same source they have descended from the same progenitor the case those species which have undergone during whole geological periods but little modification there not much difficulty believing that they may have migrated from the same region for during the vast geographical and climatal changes which will have supervened since ancient times almost any amount migration possible but many other cases which have reason believe that the species genus have been produced within comparatively recent times there great difficulty this head also obvious that the individuals the same species though now inhabiting distant and isolated regions must have proceeded from one spot where their parents were first produced for explained the last chapter incredible that individuals identically the same should ever have been produced through natural selection from parents specifically distinct are thus brought the question which has been largely discussed naturalists namely whether species have been created one more points the earth surface undoubtedly there are very many cases extreme difficulty understanding how the same species could possibly have migrated from some one point the several distant and isolated points where now found nevertheless the simplicity the view that each species was first produced within single region captivates the mind who rejects rejects the vera causa ordinary generation with subsequent migration and calls the agency miracle universally admitted that most cases the area inhabited species continuous and when plant animal inhabits two points distant from each other with interval such nature that the space could not easily passed over migration the fact given something remarkable and exceptional the capacity migrating across the sea more distinctly limited terrestrial mammals than perhaps any other organic beings and accordingly find inexplicable cases the same mammal inhabiting distant points the world geologist will feel any difficulty such cases great britain having been formerly united europe and consequently possessing the same quadrupeds but the same species can produced two separate points why not find single mammal common europe and australia south america the conditions life are nearly the same that multitude european animals and plants have become naturalised america and australia and some the aboriginal plants are identically the same these distant points the northern and southern hemispheres the answer believe that mammals have not been able migrate whereas some plants from their varied means dispersal have migrated across the vast and broken interspace the great and striking influence which barriers every kind have had distribution intelligible only the view that the great majority species have been produced one side alone and have not been able migrate the other side some few families many sub families very many genera and still greater number sections genera are confined single region and has been observed several naturalists that the most natural genera those genera which the species are most closely related each other are generally local confined one area what strange anomaly would when coming one step lower the series the individuals the same species directly opposite rule prevailed and species were not local but had been produced two more distinct areas hence seems has many other naturalists that the view each species having been produced one area alone and having subsequently migrated from that area far its powers migration and subsistence under past and present conditions permitted the most probable undoubtedly many cases occur which cannot explain how the same species could have passed from one point the other but the geographical and climatal changes which have certainly occurred within recent geological times must have interrupted rendered discontinuous the formerly continuous range many species that are reduced consider whether the exceptions continuity range are numerous and grave nature that ought give the belief rendered probable general considerations that each species has been produced within one area and has migrated thence far could would hopelessly tedious discuss all the exceptional cases the same species now living distant and separated points nor for moment pretend that any explanation could offered many such cases but after some preliminary remarks will discuss few the most striking classes facts namely the existence the same species the summits distant mountain ranges and distant points the arctic and antarctic regions and secondly the following chapter the wide distribution freshwater productions and thirdly the occurrence the same terrestrial species islands and the mainland though separated hundreds miles open sea the existence the same species distant and isolated points the earth surface can many instances explained the view each species having migrated from single birthplace then considering our ignorance with respect former climatal and geographical changes and various occasional means transport the belief that this has been the universal law seems incomparably the safest discussing this subject shall enabled the same time consider point equally important for namely whether the several distinct species genus which theory have all descended from common progenitor can have migrated undergoing modification during some part their migration from the area inhabited their progenitor can shown almost invariably the case that region which most its inhabitants are closely related belong the same genera with the species second region has probably received some former period immigrants from this other region theory will strengthened for can clearly understand the principle modification why the inhabitants region should related those another region whence has been stocked volcanic island for instance upheaved and formed the distance few hundreds miles from continent would probably receive from the course time few colonists and their descendants though modified would still plainly related inheritance the inhabitants the continent cases this nature are common and are shall hereafter more fully see inexplicable the theory independent creation this view the relation species one region those another does not differ much substituting the word variety for species from that lately advanced ingenious paper wallace which concludes that every species has come into existence coincident both space and time with pre existing closely allied species and now know from correspondence that this coincidence attributes generation with modification the previous remarks single and multiple centres creation not directly bear another allied question namely whether all the individuals the same species have descended from single pair single hermaphrodite whether some authors suppose from many individuals simultaneously created with those organic beings which never intercross such exist the species theory must have descended from succession improved varieties which will never have blended with other individuals varieties but will have supplanted each other that each successive stage modification and improvement all the individuals each variety will have descended from single parent but the majority cases namely with all organisms which habitually unite for each birth which often intercross believe that during the slow process modification the individuals the species will have been kept nearly uniform intercrossing that many individuals will have gone simultaneously changing and the whole amount modification will not have been due each stage descent from single parent illustrate what mean our english racehorses differ slightly from the horses every other breed but they not owe their difference and superiority descent from any single pair but continued care selecting and training many individuals during many generations before discussing the three classes facts which have selected presenting the greatest amount difficulty the theory single centres creation must say few words the means dispersal means dispersal sir lyell and other authors have ably treated this subject can give here only the briefest abstract the more important facts change climate must have had powerful influence migration region when its climate was different may have been high road for migration but now impassable shall however presently have discuss this branch the subject some detail changes level the land must also have been highly influential narrow isthmus now separates two marine faunas submerge let formerly have been submerged and the two faunas will now blend may formerly have blended where the sea now extends land may former period have connected islands possibly even continents together and thus have allowed terrestrial productions pass from one the other geologist will dispute that great mutations level have occurred within the period existing organisms edward forbes insisted that all the islands the atlantic must recently have been connected with europe africa and europe likewise with america other authors have thus hypothetically bridged over every ocean and have united almost every island some mainland indeed the arguments used forbes are trusted must admitted that scarcely single island exists which has not recently been united some continent this view cuts the gordian knot the dispersal the same species the most distant points and removes many difficulty but the best judgment are not authorized admitting such enormous geographical changes within the period existing species seems that have abundant evidence great oscillations level our continents but not such vast changes their position and extension have united them within the recent period each other and the several intervening oceanic islands freely admit the former existence many islands now buried beneath the sea which may have served halting places for plants and for many animals during their migration the coral producing oceans such sunken islands are now marked believe rings coral atolls standing over them whenever fully admitted believe will some day that each species has proceeded from single birthplace and when the course time know something definite about the means distribution shall enabled speculate with security the former extension the land but not believe that will ever proved that within the recent period continents which are now quite separate have been continuously almost continuously united with each other and with the many existing oceanic islands several facts distribution such the great difference the marine faunas the opposite sides almost every continent the close relation the tertiary inhabitants several lands and even seas their present inhabitants certain degree relation shall hereafter see between the distribution mammals and the depth the sea these and other such facts seem opposed the admission such prodigious geographical revolutions within the recent period are necessitated the view advanced forbes and admitted his many followers the nature and relative proportions the inhabitants oceanic islands likewise seem opposed the belief their former continuity with continents nor does their almost universally volcanic composition favour the admission that they are the wrecks sunken continents they had originally existed mountain ranges the land some least the islands would have been formed like other mountain summits granite metamorphic schists old fossiliferous other such rocks instead consisting mere piles volcanic matter must now say few words what are called accidental means but which more properly might called occasional means distribution shall here confine myself plants botanical works this that plant stated ill adapted for wide dissemination but for transport across the sea the greater less facilities may said almost wholly unknown until tried with berkeley aid few experiments was not even known how far seeds could resist the injurious action sea water surprise found that out kinds germinated after immersion days and few survived immersion 137 days for convenience sake chiefly tried small seeds without the capsule fruit and all these sank few days they could not floated across wide spaces the sea whether not they were injured the salt water afterwards tried some larger fruits capsules etc and some these floated for long time well known what difference there the buoyancy green and seasoned timber and occurred that floods might wash down plants branches and that these might dried the banks and then fresh rise the stream washed into the sea hence was led dry stems and branches plants with ripe fruit and place them sea water the majority sank quickly but some which whilst green floated for very short time when dried floated much longer for instance ripe hazel nuts sank immediately but when dried they floated for days and afterwards when planted they germinated asparagus plant with ripe berries floated for days when dried floated for days and the seeds afterwards germinated the ripe seeds helosciadium sank two days when dried they floated for above days and afterwards germinated altogether out the dried plants floated for above days and some the floated for very much longer period that seeds germinated after immersion days and plants with ripe fruit but not all the same species the foregoing experiment floated after being dried for above days far may infer anything from these scanty facts may conclude that the seeds 100 plants any country might floated sea currents during days and would retain their power germination johnston physical atlas the average rate the several atlantic currents miles per diem some currents running the rate miles per diem this average the seeds 100 plants belonging one country might floated across 924 miles sea another country and when stranded blown favourable spot inland gale they would germinate subsequently experiments martens tried similar ones but much better manner for placed the seeds box the actual sea that they were alternately wet and exposed the air like really floating plants tried seeds mostly different from mine but chose many large fruits and likewise seeds from plants which live near the sea and this would have favoured the average length their flotation and their resistance the injurious action the salt water the other hand did not previously dry the plants branches with the fruit and this have seen would have caused some them have floated much longer the result was that his seeds floated for days and were then capable germination but not doubt that plants exposed the waves would float for less time than those protected from violent movement our experiments therefore would perhaps safer assume that the seeds about 100 plants flora after having been dried could floated across space sea 900 miles width and would then germinate the fact the larger fruits often floating longer than the small interesting plants with large seeds fruit could hardly transported any other means and alph candolle has shown that such plants generally have restricted ranges but seeds may occasionally transported another manner drift timber thrown most islands even those the midst the widest oceans and the natives the coral islands the pacific procure stones for their tools solely from the roots drifted trees these stones being valuable royal tax find examination that when irregularly shaped stones are embedded the roots trees small parcels earth are very frequently enclosed their interstices and behind them perfectly that not particle could washed away the longest transport out one small portion earth thus completely enclosed wood oak about years old three dicotyledonous plants germinated certain the accuracy this observation again can show that the carcasses birds when floating the sea sometimes escape being immediately devoured and seeds many kinds the crops floating birds long retain their vitality peas and vetches for instance are killed even few days immersion sea water but some taken out the crop pigeon which had floated artificial salt water for days surprise nearly all germinated living birds can hardly fail highly effective agents the transportation seeds could give many facts showing how frequently birds many kinds are blown gales vast distances across the ocean may think safely assume that under such circumstances their rate flight would often miles hour and some authors have given far higher estimate have never seen instance nutritious seeds passing through the intestines bird but hard seeds fruit will pass uninjured through even the digestive organs turkey the course two months picked garden kinds seeds out the excrement small birds and these seemed perfect and some them which tried germinated but the following fact more important the crops birds not secrete gastric juice and not the least injure know trial the germination seeds now after bird has found and devoured large supply food positively asserted that all the grains not pass into the gizzard for even hours bird this interval might easily blown the distance 500 miles and hawks are known look out for tired birds and the contents their torn crops might thus readily get scattered brent informs that friend his had give flying carrier pigeons from france england the hawks the english coast destroyed many their arrival some hawks and owls bolt their prey whole and after interval from twelve twenty hours disgorge pellets which know from experiments made the zoological gardens include seeds capable germination some seeds the oat wheat millet canary hemp clover and beet germinated after having been from twelve twenty one hours the stomachs different birds prey and two seeds beet grew after having been thus retained for two days and fourteen hours freshwater fish find eat seeds many land and water plants fish are frequently devoured birds and thus the seeds might transported from place place forced many kinds seeds into the stomachs dead fish and then gave their bodies fishing eagles storks and pelicans these birds after interval many hours either rejected the seeds pellets passed them their excrement and several these seeds retained their power germination certain seeds however were always killed this process although the beaks and feet birds are generally quite clean can show that earth sometimes adheres them one instance removed twenty two grains dry argillaceous earth from one foot partridge and this earth there was pebble quite large the seed vetch thus seeds might occasionally transported great distances for many facts could given showing that soil almost everywhere charged with seeds reflect for moment the millions quails which annually cross the mediterranean and can doubt that the earth adhering their feet would sometimes include few minute seeds but shall presently have recur this subject icebergs are known sometimes loaded with earth and stones and have even carried brushwood bones and the nest land bird can hardly doubt that they must occasionally have transported seeds from one part another the arctic and antarctic regions suggested lyell and during the glacial period from one part the now temperate regions another the azores from the large number the species plants common europe comparison with the plants other oceanic islands nearer the mainland and remarked watson from the somewhat northern character the flora comparison with the latitude suspected that these islands had been partly stocked ice borne seeds during the glacial epoch request sir lyell wrote hartung inquire whether had observed erratic boulders these islands and answered that had found large fragments granite and other rocks which not occur the archipelago hence may safely infer that icebergs formerly landed their rocky burthens the shores these mid ocean islands and least possible that they may have brought thither the seeds northern plants considering that the several above means transport and that several other means which without doubt remain discovered have been action year after year for centuries and tens thousands years would think marvellous fact many plants had not thus become widely transported these means transport are sometimes called accidental but this not strictly correct the currents the sea are not accidental nor the direction prevalent gales wind should observed that scarcely any means transport would carry seeds for very great distances for seeds not retain their vitality when exposed for great length time the action seawater nor could they long carried the crops intestines birds these means however would suffice for occasional transport across tracts sea some hundred miles breadth from island island from continent neighbouring island but not from one distant continent another the floras distant continents would not such means become mingled any great degree but would remain distinct now see them the currents from their course would never bring seeds from north america britain though they might and bring seeds from the west indies our western shores where not killed long immersion salt water they could not endure our climate almost every year one two land birds are blown across the whole atlantic ocean from north america the western shores ireland and england but seeds could transported these wanderers only one means namely dirt sticking their feet which itself rare accident even this case how small would the chance seed falling favourable soil and coming maturity but would great error argue that because well stocked island like great britain has not far known and would very difficult prove this received within the last few centuries through occasional means transport immigrants from europe any other continent that poorly stocked island though standing more remote from the mainland would not receive colonists similar means not doubt that out twenty seeds animals transported island even far less well stocked than britain scarcely more than one would well fitted its new home become naturalised but this seems valid argument against what would effected occasional means transport during the long lapse geological time whilst island was being upheaved and formed and before had become fully stocked with inhabitants almost bare land with few destructive insects birds living there nearly every seed which chanced arrive would sure germinate and survive dispersal during the glacial period the identity many plants and animals mountain summits separated from each other hundreds miles lowlands where the alpine species could not possibly exist one the most striking cases known the same species living distant points without the apparent possibility their having migrated from one the other indeed remarkable fact see many the same plants living the snowy regions the alps pyrenees and the extreme northern parts europe but far more remarkable that the plants the white mountains the united states america are all the same with those labrador and nearly all the same hear from asa gray with those the loftiest mountains europe even long ago 1747 such facts led gmelin conclude that the same species must have been independently created several distinct points and might have remained this same belief had not agassiz and others called vivid attention the glacial period which shall immediately see affords simple explanation these facts have evidence almost every conceivable kind organic and inorganic that within very recent geological period central europe and north america suffered under arctic climate the ruins house burnt fire not tell their tale more plainly than the mountains scotland and wales with their scored flanks polished surfaces and perched boulders the icy streams with which their valleys were lately filled greatly has the climate europe changed that northern italy gigantic moraines left old glaciers are now clothed the vine and maize throughout large part the united states erratic boulders and rocks scored drifted icebergs and coast ice plainly reveal former cold period the former influence the glacial climate the distribution the inhabitants europe explained with remarkable clearness edward forbes substantially follows but shall follow the changes more readily supposing new glacial period come slowly and then pass away formerly occurred the cold came and each more southern zone became fitted for arctic beings and ill fitted for their former more temperate inhabitants the latter would supplanted and arctic productions would take their places the inhabitants the more temperate regions would the same time travel southward unless they were stopped barriers which case they would perish the mountains would become covered with snow and ice and their former alpine inhabitants would descend the plains the time that the cold had reached its maximum should have uniform arctic fauna and flora covering the central parts europe far south the alps and pyrenees and even stretching into spain the now temperate regions the united states would likewise covered arctic plants and animals and these would nearly the same with those europe for the present circumpolar inhabitants which suppose have everywhere travelled southward are remarkably uniform round the world may suppose that the glacial period came little earlier later north america than europe will the southern migration there have been little earlier later but this will make difference the final result the warmth returned the arctic forms would retreat northward closely followed their retreat the productions the more temperate regions and the snow melted from the bases the mountains the arctic forms would seize the cleared and thawed ground always ascending higher and higher the warmth increased whilst their brethren were pursuing their northern journey hence when the warmth had fully returned the same arctic species which had lately lived body together the lowlands the old and new worlds would left isolated distant mountain summits having been exterminated all lesser heights and the arctic regions both hemispheres thus can understand the identity many plants points immensely remote the mountains the united states and europe can thus also understand the fact that the alpine plants each mountain range are more especially related the arctic forms living due north nearly due north them for the migration the cold came and the migration the returning warmth will generally have been due south and north the alpine plants for example scotland remarked watson and those the pyrenees remarked ramond are more especially allied the plants northern scandinavia those the united states labrador those the mountains siberia the arctic regions that country these views grounded they are the perfectly well ascertained occurrence former glacial period seem explain satisfactory manner the present distribution the alpine and arctic productions europe and america that when other regions find the same species distant mountain summits may almost conclude without other evidence that colder climate permitted their former migration across the low intervening tracts since become too warm for their existence the climate since the glacial period has ever been any degree warmer than present some geologists the united states believe have been the case chiefly from the distribution the fossil gnathodon then the arctic and temperate productions will very late period have marched little further north and subsequently have retreated their present homes but have met with satisfactory evidence with respect this intercalated slightly warmer period since the glacial period the arctic forms during their long southern migration and migration northward will have been exposed nearly the same climate and especially noticed they will have kept body together consequently their mutual relations will not have been much disturbed and accordance with the principles inculcated this volume they will not have been liable much modification but with our alpine productions left isolated from the moment the returning warmth first the bases and ultimately the summits the mountains the case will have been somewhat different for not likely that all the same arctic species will have been left mountain ranges distant from each other and have survived there ever since they will also all probability have become mingled with ancient alpine species which must have existed the mountains before the commencement the glacial epoch and which during its coldest period will have been temporarily driven down the plains they will also have been exposed somewhat different climatal influences their mutual relations will thus have been some degree disturbed consequently they will have been liable modification and this find has been the case for compare the present alpine plants and animals the several great european mountain ranges though very many the species are identically the same some present varieties some are ranked doubtful forms and some few are distinct yet closely allied representative species illustrating what believe actually took place during the glacial period assumed that its commencement the arctic productions were uniform round the polar regions they are the present day but the foregoing remarks distribution apply not only strictly arctic forms but also many sub arctic and some few northern temperate forms for some these are the same the lower mountains and the plains north america and europe and may reasonably asked how account for the necessary degree uniformity the sub arctic and northern temperate forms round the world the commencement the glacial period the present day the sub arctic and northern temperate productions the old and new worlds are separated from each other the atlantic ocean and the extreme northern part the pacific during the glacial period when the inhabitants the old and new worlds lived further southwards than present they must have been still more completely separated wider spaces ocean believe the above difficulty may surmounted looking still earlier changes climate opposite nature have good reason believe that during the newer pliocene period before the glacial epoch and whilst the majority the inhabitants the world were specifically the same now the climate was warmer than the present day hence may suppose that the organisms now living under the climate latitude during the pliocene period lived further north under the polar circle latitude and that the strictly arctic productions then lived the broken land still nearer the pole now look globe shall see that under the polar circle there almost continuous land from western europe through siberia eastern america and this continuity the circumpolar land and the consequent freedom for intermigration under more favourable climate attribute the necessary amount uniformity the sub arctic and northern temperate productions the old and new worlds period anterior the glacial epoch believing from reasons before alluded that our continents have long remained nearly the same relative position though subjected large but partial oscillations level strongly inclined extend the above view and infer that during some earlier and still warmer period such the older pliocene period large number the same plants and animals inhabited the almost continuous circumpolar land and that these plants and animals both the old and new worlds began slowly migrate southwards the climate became less warm long before the commencement the glacial period now see believe their descendants mostly modified condition the central parts europe and the united states this view can understand the relationship with very little identity between the productions north america and europe relationship which most remarkable considering the distance the two areas and their separation the atlantic ocean can further understand the singular fact remarked several observers that the productions europe and america during the later tertiary stages were more closely related each other than they are the present time for during these warmer periods the northern parts the old and new worlds will have been almost continuously united land serving bridge since rendered impassable cold for the inter migration their inhabitants during the slowly decreasing warmth the pliocene period soon the species common which inhabited the new and old worlds migrated south the polar circle they must have been completely cut off from each other this separation far the more temperate productions are concerned took place long ages ago and the plants and animals migrated southward they will have become mingled the one great region with the native american productions and have had compete with them and the other great region with those the old world consequently have here everything favourable for much modification for far more modification than with the alpine productions left isolated within much more recent period the several mountain ranges and the arctic lands the two worlds hence has come that when compare the now living productions the temperate regions the new and old worlds find very few identical species though asa gray has lately shown that more plants are identical than was formerly supposed but find every great class many forms which some naturalists rank geographical races and others distinct species and host closely allied representative forms which are ranked all naturalists specifically distinct the land the waters the sea slow southern migration marine fauna which during the pliocene even somewhat earlier period was nearly uniform along the continuous shores the polar circle will account the theory modification for many closely allied forms now living areas completely sundered thus think can understand the presence many existing and tertiary representative forms the eastern and western shores temperate north america and the still more striking case many closely allied crustaceans described dana admirable work some fish and other marine animals the mediterranean and the seas japan areas now separated continent and nearly hemisphere equatorial ocean these cases relationship without identity the inhabitants seas now disjoined and likewise the past and present inhabitants the temperate lands north america and europe are inexplicable the theory creation cannot say that they have been created alike correspondence with the nearly similar physical conditions the areas for compare for instance certain parts south america with the southern continents the old world see countries closely corresponding all their physical conditions but with their inhabitants utterly dissimilar but must return our more immediate subject the glacial period convinced that forbes view may largely extended europe have the plainest evidence the cold period from the western shores britain the oural range and southward the pyrenees may infer from the frozen mammals and nature the mountain vegetation that siberia was similarly affected along the himalaya points 900 miles apart glaciers have left the marks their former low descent and sikkim hooker saw maize growing gigantic ancient moraines south the equator have some direct evidence former glacial action new zealand and the same plants found widely separated mountains this island tell the same story one account which has been published can trusted have direct evidence glacial action the south eastern corner australia looking america the northern half ice borne fragments rock have been observed the eastern side far south lat and the shores the pacific where the climate now different far south lat deg erratic boulders have also been noticed the rocky mountains the cordillera equatorial south america glaciers once extended far below their present level central chile was astonished the structure vast mound detritus about 800 feet height crossing valley the andes and this now feel convinced was gigantic moraine left far below any existing glacier further south both sides the continent from lat the southernmost extremity have the clearest evidence former glacial action huge boulders transported far from their parent source not know that the glacial epoch was strictly simultaneous these several far distant points opposite sides the world but have good evidence almost every case that the epoch was included within the latest geological period have also excellent evidence that endured for enormous time measured years each point the cold may have come have ceased earlier one point the globe than another but seeing that endured for long each and that was contemporaneous geological sense seems probable that was during part least the period actually simultaneous throughout the world without some distinct evidence the contrary may least admit probable that the glacial action was simultaneous the eastern and western sides north america the cordillera under the equator and under the warmer temperate zones and both sides the southern extremity the continent this admitted difficult avoid believing that the temperature the whole world was this period simultaneously cooler but would suffice for purpose the temperature was the same time lower along certain broad belts longitude this view the whole world least broad longitudinal belts having been simultaneously colder from pole pole much light can thrown the present distribution identical and allied species america hooker has shown that between forty and fifty the flowering plants tierra del fuego forming inconsiderable part its scanty flora are common europe enormously remote these two points are and there are many closely allied species the lofty mountains equatorial america host peculiar species belonging european genera occur the highest mountains brazil some few european genera were found gardner which not exist the wide intervening hot countries the silla caraccas the illustrious humboldt long ago found species belonging genera characteristic the cordillera the mountains abyssinia several european forms and some few representatives the peculiar flora the cape good hope occur the cape good hope very few european species believed not have been introduced man and the mountains some few representative european forms are found which have not been discovered the intertropical parts africa the himalaya and the isolated mountain ranges the peninsula india the heights ceylon and the volcanic cones java many plants occur either identically the same representing each other and the same time representing plants europe not found the intervening hot lowlands list the genera collected the loftier peaks java raises picture collection made hill europe still more striking the fact that southern australian forms are clearly represented plants growing the summits the mountains borneo some these australian forms hear from hooker extend along the heights the peninsula malacca and are thinly scattered the one hand over india and the other far north japan the southern mountains australia müller has discovered several european species other species not introduced man occur the lowlands and long list can given informed hooker european genera found australia but not the intermediate torrid regions the admirable introduction the flora new zealand hooker analogous and striking facts are given regard the plants that large island hence see that throughout the world the plants growing the more lofty mountains and the temperate lowlands the northern and southern hemispheres are sometimes identically the same but they are much oftener specifically distinct though related each other most remarkable manner this brief abstract applies plants alone some strictly analogous facts could given the distribution terrestrial animals marine productions similar cases occur example may quote remark the highest authority professor dana that certainly wonderful fact that new zealand should have closer resemblance its crustacea great britain its antipode than any other part the world sir richardson also speaks the reappearance the shores new zealand tasmania etc northern forms fish hooker informs that twenty five species algæ are common new zealand and europe but have not been found the intermediate tropical seas should observed that the northern species and forms found the southern parts the southern hemisphere and the mountain ranges the intertropical regions are not arctic but belong the northern temperate zones watson has recently remarked receding from polar towards equatorial latitudes the alpine mountain floras really become less and less arctic many the forms living the mountains the warmer regions the earth and the southern hemisphere are doubtful value being ranked some naturalists specifically distinct others varieties but some are certainly identical and many though closely related northern forms must ranked distinct species now let see what light can thrown the foregoing facts the belief supported large body geological evidence that the whole world large part was during the glacial period simultaneously much colder than present the glacial period measured years must have been very long and when remember over what vast spaces some naturalised plants and animals have spread within few centuries this period will have been ample for any amount migration the cold came slowly all the tropical plants and other productions will have retreated from both sides towards the equator followed the rear the temperate productions and these the arctic but with the latter are not now concerned the tropical plants probably suffered much extinction how much one can say perhaps formerly the tropics supported many species see the present day crowded together the cape good hope and parts temperate australia know that many tropical plants and animals can withstand considerable amount cold many might have escaped extermination during moderate fall temperature more especially escaping into the warmest spots but the great fact bear mind that all tropical productions will have suffered certain extent the other hand the temperate productions after migrating nearer the equator though they will have been placed under somewhat new conditions will have suffered less and certain that many temperate plants protected from the inroads competitors can withstand much warmer climate than their own hence seems possible bearing mind that the tropical productions were suffering state and could not have presented firm front against intruders that certain number the more vigorous and dominant temperate forms might have penetrated the native ranks and have reached even crossed the equator the invasion would course have been greatly favoured high land and perhaps dry climate for falconer informs that the damp with the heat the tropics which destructive perennial plants from temperate climate the other hand the most humid and hottest districts will have afforded asylum the tropical natives the mountain ranges north west the himalaya and the long line the cordillera seem have afforded two great lines invasion and striking fact lately communicated hooker that all the flowering plants about forty six number common tierra del fuego and europe still exist north america which must have lain the line march but not doubt that some temperate productions entered and crossed even the lowlands the tropics the period when the cold was most intense when arctic forms had migrated some twenty five degrees latitude from their native country and covered the land the foot the pyrenees this period extreme cold believe that the climate under the equator the level the sea was about the same with that now felt there the height six seven thousand feet during this the coldest period suppose that large spaces the tropical lowlands were clothed with mingled tropical and temperate vegetation like that now growing with strange luxuriance the base the himalaya graphically described hooker thus believe considerable number plants few terrestrial animals and some marine productions migrated during the glacial period from the northern and southern temperate zones into the intertropical regions and some even crossed the equator the warmth returned these temperate forms would naturally ascend the higher mountains being exterminated the lowlands those which had not reached the equator would migrate northward southward towards their former homes but the forms chiefly northern which had crossed the equator would travel still further from their homes into the more temperate latitudes the opposite hemisphere although have reason believe from geological evidence that the whole body arctic shells underwent scarcely any modification during their long southern migration and migration northward the case may have been wholly different with those intruding forms which settled themselves the intertropical mountains and the southern hemisphere these being surrounded strangers will have had compete with many new forms life and probable that selected modifications their structure habits and constitutions will have profited them thus many these wanderers though still plainly related inheritance their brethren the northern southern hemispheres now exist their new homes well marked varieties distinct species remarkable fact strongly insisted hooker regard america and alph candolle regard australia that many more identical plants and allied forms have apparently migrated from the north the south than reversed direction see however few southern vegetable forms the mountains borneo and abyssinia suspect that this preponderant migration from north south due the greater extent land the north and the northern forms having existed their own homes greater numbers and having consequently been advanced through natural selection and competition higher stage perfection dominating power than the southern forms and thus when they became commingled during the glacial period the northern forms were enabled beat the less powerful southern forms just the same manner see the present day that very many european productions cover the ground plata and lesser degree australia and have certain extent beaten the natives whereas extremely few southern forms have become naturalised any part europe though hides wool and other objects likely carry seeds have been largely imported into europe during the last two three centuries from plata and during the last thirty forty years from australia something the same kind must have occurred the intertropical mountains doubt before the glacial period they were stocked with endemic alpine forms but these have almost everywhere largely yielded the more dominant forms generated the larger areas and more efficient workshops the north many islands the native productions are nearly equalled even outnumbered the naturalised and the natives have not been actually exterminated their numbers have been greatly reduced and this the first stage towards extinction mountain island the land and the intertropical mountains before the glacial period must have been completely isolated and believe that the productions these islands the land yielded those produced within the larger areas the north just the same way the productions real islands have everywhere lately yielded continental forms naturalised man agency far from supposing that all difficulties are removed the view here given regard the range and affinities the allied species which live the northern and southern temperate zones and the mountains the intertropical regions very many difficulties remain solved not pretend indicate the exact lines and means migration the reason why certain species and not others have migrated why certain species have been modified and have given rise new groups forms and others have remained unaltered cannot hope explain such facts until can say why one species and not another becomes naturalised man agency foreign land why one ranges twice thrice far and twice thrice common another species within their own homes have said that many difficulties remain solved some the most remarkable are stated with admirable clearness hooker his botanical works the antarctic regions these cannot here discussed will only say that far regards the occurrence identical species points enormously remote kerguelen land new zealand and fuegia believe that towards the close the glacial period icebergs suggested lyell have been largely concerned their dispersal but the existence several quite distinct species belonging genera exclusively confined the south these and other distant points the southern hemisphere theory descent with modification far more remarkable case difficulty for some these species are distinct that cannot suppose that there has been time since the commencement the glacial period for their migration and for their subsequent modification the necessary degree the facts seem indicate that peculiar and very distinct species have migrated radiating lines from some common centre and inclined look the southern the northern hemisphere former and warmer period before the commencement the glacial period when the antarctic lands now covered with ice supported highly peculiar and isolated flora suspect that before this flora was exterminated the glacial epoch few forms were widely dispersed various points the southern hemisphere occasional means transport and the aid halting places existing and now sunken islands and perhaps the commencement the glacial period icebergs these means believe the southern shores america australia new zealand have become slightly tinted the same peculiar forms vegetable life sir lyell striking passage has speculated language almost identical with mine the effects great alternations climate geographical distribution believe that the world has recently felt one his great cycles change and that this view combined with modification through natural selection multitude facts the present distribution both the same and allied forms life can explained the living waters may said have flowed during one short period from the north and from the south and have crossed the equator but have flowed with greater force from the north have freely inundated the south the tide leaves its drift horizontal lines though rising higher the shores where the tide rises highest have the living waters left their living drift our mountain summits line gently rising from the arctic lowlands great height under the equator the various beings thus left stranded may compared with savage races man driven and surviving the mountain fastnesses almost every land which serve record full interest the former inhabitants the surrounding lowlands chapter xii geographical distribution continued distribution fresh water productions the inhabitants oceanic islands absence batrachians and terrestrial mammals the relation the inhabitants islands those the nearest mainland colonisation from the nearest source with subsequent modification summary the last and present chapters lakes and river systems are separated from each other barriers land might have been thought that fresh water productions would not have ranged widely within the same country and the sea apparently still more impassable barrier that they never would have extended distant countries but the case exactly the reverse not only have many fresh water species belonging quite different classes enormous range but allied species prevail remarkable manner throughout the world well remember when first collecting the fresh waters brazil feeling much surprise the similarity the fresh water insects shells etc and the dissimilarity the surrounding terrestrial beings compared with those britain but this power fresh water productions ranging widely though unexpected can think most cases explained their having become fitted manner highly useful them for short and frequent migrations from pond pond from stream stream and liability wide dispersal would follow from this capacity almost necessary consequence can here consider only few cases regard fish believe that the same species never occur the fresh waters distant continents but the same continent the species often range widely and almost capriciously for two river systems will have some fish common and some different few facts seem favour the possibility their occasional transport accidental means like that the live fish not rarely dropped whirlwinds india and the vitality their ova when removed from the water but inclined attribute the dispersal fresh water fish mainly slight changes within the recent period the level the land having caused rivers flow into each other instances also could given this having occurred during floods without any change level have evidence the loess the rhine considerable changes level the land within very recent geological period and when the surface was peopled existing land and fresh water shells the wide difference the fish opposite sides continuous mountain ranges which from early period must have parted river systems and completely prevented their inosculation seems lead this same conclusion with respect allied fresh water fish occurring very distant points the world doubt there are many cases which cannot present explained but some fresh water fish belong very ancient forms and such cases there will have been ample time for great geographical changes and consequently time and means for much migration the second place salt water fish can with care slowly accustomed live fresh water and according valenciennes there hardly single group fishes confined exclusively fresh water that may imagine that marine member fresh water group might travel far along the shores the sea and subsequently become modified and adapted the fresh waters distant land some species fresh water shells have very wide range and allied species which theory are descended from common parent and must have proceeded from single source prevail throughout the world their distribution first perplexed much their ova are not likely transported birds and they are immediately killed sea water are the adults could not even understand how some naturalised species have rapidly spread throughout the same country but two facts which have observed and doubt many others remain observed throw some light this subject when duck suddenly emerges from pond covered with duck weed have twice seen these little plants adhering its back and has happened removing little duck weed from one aquarium another that have quite unintentionally stocked the one with fresh water shells from the other but another agency perhaps more effectual suspended duck feet which might represent those bird sleeping natural pond aquarium where many ova fresh water shells were hatching and found that numbers the extremely minute and just hatched shells crawled the feet and clung them firmly that when taken out the water they could not jarred off though somewhat more advanced age they would voluntarily drop off these just hatched molluscs though aquatic their nature survived the duck feet damp air from twelve twenty hours and this length time duck heron might fly least six seven hundred miles and would sure alight pool rivulet blown across sea oceanic island any other distant point sir charles lyell also informs that dyticus has been caught with ancylus fresh water shell like limpet firmly adhering and water beetle the same family colymbetes once flew board the beagle when forty five miles distant from the nearest land how much farther might have flown with favouring gale one can tell with respect plants has long been known what enormous ranges many fresh water and even marsh species have both over continents and the most remote oceanic islands this strikingly shown remarked alph candolle large groups terrestrial plants which have only very few aquatic members for these latter seem immediately acquire consequence very wide range think favourable means dispersal explain this fact have before mentioned that earth occasionally though rarely adheres some quantity the feet and beaks birds wading birds which frequent the muddy edges ponds suddenly flushed would the most likely have muddy feet birds this order can show are the greatest wanderers and are occasionally found the most remote and barren islands the open ocean they would not likely alight the surface the sea that the dirt would not washed off their feet when making land they would sure fly their natural fresh water haunts not believe that botanists are aware how charged the mud ponds with seeds have tried several little experiments but will here give only the most striking case took february three table spoonfuls mud from three different points beneath water the edge little pond this mud when dry weighed only ounces kept covered study for six months pulling and counting each plant grew the plants were many kinds and were altogether 537 number and yet the viscid mud was all contained breakfast cup considering these facts think would inexplicable circumstance water birds did not transport the seeds fresh water plants vast distances and consequently the range these plants was not very great the same agency may have come into play with the eggs some the smaller fresh water animals other and unknown agencies probably have also played part have stated that fresh water fish eat some kinds seeds though they reject many other kinds after having swallowed them even small fish swallow seeds moderate size the yellow water lily and potamogeton herons and other birds century after century have gone daily devouring fish they then take flight and other waters are blown across the sea and have seen that seeds retain their power germination when rejected pellets excrement many hours afterwards when saw the great size the seeds that fine water lily the nelumbium and remembered alph candolle remarks this plant thought that its distribution must remain quite inexplicable but audubon states that found the seeds the great southern water lily probably according hooker the nelumbium luteum heron stomach although not know the fact yet analogy makes believe that heron flying another pond and getting hearty meal fish would probably reject from its stomach pellet containing the seeds the nelumbium undigested the seeds might dropped the bird whilst feeding its young the same way fish are known sometimes dropped considering these several means distribution should remembered that when pond stream first formed for instance rising islet will unoccupied and single seed egg will have good chance succeeding although there will always struggle for life between the individuals the species however few already occupying any pond yet the number kinds small compared with those the land the competition will probably less severe between aquatic than between terrestrial species consequently intruder from the waters foreign country would have better chance seizing place than the case terrestrial colonists should also remember that some perhaps many fresh water productions are low the scale nature and that have reason believe that such low beings change become modified less quickly than the high and this will give longer time than the average for the migration the same aquatic species should not forget the probability many species having formerly ranged continuously fresh water productions ever can range over immense areas and having subsequently become extinct intermediate regions but the wide distribution fresh water plants and the lower animals whether retaining the same identical form some degree modified believe mainly depends the wide dispersal their seeds and eggs animals more especially fresh water birds which have large powers flight and naturally travel from one another and often distant piece water nature like careful gardener thus takes her seeds from bed particular nature and drops them another equally well fitted for them the inhabitants oceanic islands now come the last the three classes facts which have selected presenting the greatest amount difficulty the view that all the individuals both the same and allied species have descended from single parent and therefore have all proceeded from common birthplace notwithstanding that the course time they have come inhabit distant points the globe have already stated that cannot honestly admit forbes view continental extensions which legitimately followed out would lead the belief that within the recent period all existing islands have been nearly quite joined some continent this view would remove many difficulties but would not think explain all the facts regard insular productions the following remarks shall not confine myself the mere question dispersal but shall consider some other facts which bear the truth the two theories independent creation and descent with modification the species all kinds which inhabit oceanic islands are few number compared with those equal continental areas alph candolle admits this for plants and wollaston for insects look the large size and varied stations new zealand extending over 780 miles latitude and compare its flowering plants only 750 number with those equal area the cape good hope australia must think admit that something quite independently any difference physical conditions has caused great difference number even the uniform county cambridge has 847 plants and the little island anglesea 764 but few ferns and few introduced plants are included these numbers and the comparison some other respects not quite fair have evidence that the barren island ascension aboriginally possessed under half dozen flowering plants yet many have become naturalised they have new zealand and every other oceanic island which can named helena there reason believe that the naturalised plants and animals have nearly quite exterminated many native productions who admits the doctrine the creation each separate species will have admit that sufficient number the best adapted plants and animals have not been created oceanic islands for man has unintentionally stocked them from various sources far more fully and perfectly than has nature although oceanic islands the number kinds inhabitants scanty the proportion endemic species those found nowhere else the world often extremely large compare for instance the number the endemic land shells madeira the endemic birds the galapagos archipelago with the number found any continent and then compare the area the islands with that the continent shall see that this true this fact might have been expected theory for already explained species occasionally arriving after long intervals new and isolated district and having compete with new associates will eminently liable modification and will often produce groups modified descendants but means follows that because island nearly all the species one class are peculiar those another class another section the same class are peculiar and this difference seems depend the species which not become modified having immigrated with facility and body that their mutual relations have not been much disturbed thus the galapagos islands nearly every land bird but only two out the eleven marine birds are peculiar and obvious that marine birds could arrive these islands more easily than land birds bermuda the other hand which lies about the same distance from north america the galapagos islands from south america and which has very peculiar soil does not possess one endemic land bird and know from jones admirable account bermuda that very many north american birds during their great annual migrations visit either periodically occasionally this island madeira does not possess one peculiar bird and many european and african birds are almost every year blown there informed harcourt that these two islands bermuda and madeira have been stocked birds which for long ages have struggled together their former homes and have become mutually adapted each other and when settled their new homes each kind will have been kept the others their proper places and habits and will consequently have been little liable modification madeira again inhabited wonderful number peculiar land shells whereas not one species sea shell confined its shores now though not know how seashells are dispersed yet can see that their eggs larvæ perhaps attached seaweed floating timber the feet wading birds might transported far more easily than land shells across three four hundred miles open sea the different orders insects madeira apparently present analogous facts oceanic islands are sometimes deficient certain classes and their places are apparently occupied the other inhabitants the galapagos islands reptiles and new zealand gigantic wingless birds take the place mammals the plants the galapagos islands hooker has shown that the proportional numbers the different orders are very different from what they are elsewhere such cases are generally accounted for the physical conditions the islands but this explanation seems not little doubtful facility immigration believe has been least important the nature the conditions many remarkable little facts could given with respect the inhabitants remote islands for instance certain islands not tenanted mammals some the endemic plants have beautifully hooked seeds yet few relations are more striking than the adaptation hooked seeds for transportal the wool and fur quadrupeds this case presents difficulty view for hooked seed might transported island some other means and the plant then becoming slightly modified but still retaining its hooked seeds would form endemic species having useless appendage any rudimentary organ for instance the shrivelled wings under the soldered elytra many insular beetles again islands often possess trees bushes belonging orders which elsewhere include only herbaceous species now trees alph candolle has shown generally have whatever the cause may confined ranges hence trees would little likely reach distant oceanic islands and herbaceous plant though would have chance successfully competing stature with fully developed tree when established island and having compete with herbaceous plants alone might readily gain advantage growing taller and taller and overtopping the other plants natural selection would often tend add the stature herbaceous plants when growing island whatever order they belonged and thus convert them first into bushes and ultimately into trees with respect the absence whole orders oceanic islands bory vincent long ago remarked that batrachians frogs toads newts have never been found any the many islands with which the great oceans are studded have taken pains verify this assertion and have found strictly true have however been assured that frog exists the mountains the great island new zealand but suspect that this exception the information correct may explained through glacial agency this general absence frogs toads and newts many oceanic islands cannot accounted for their physical conditions indeed seems that islands are peculiarly well fitted for these animals for frogs have been introduced into madeira the azores and mauritius and have multiplied become nuisance but these animals and their spawn are known immediately killed sea water view can see that there would great difficulty their transportal across the sea and therefore why they not exist any oceanic island but why the theory creation they should not have been created there would very difficult explain mammals offer another and similar case have carefully searched the oldest voyages but have not finished search yet have not found single instance free from doubt terrestrial mammal excluding domesticated animals kept the natives inhabiting island situated above 300 miles from continent great continental island and many islands situated much less distance are equally barren the falkland islands which are inhabited wolf like fox come nearest exception but this group cannot considered oceanic lies bank connected with the mainland moreover icebergs formerly brought boulders its western shores and they may have formerly transported foxes frequently now happens the arctic regions yet cannot said that small islands will not support small mammals for they occur many parts the world very small islands close continent and hardly island can named which our smaller quadrupeds have not become naturalised and greatly multiplied cannot said the ordinary view creation that there has not been time for the creation mammals many volcanic islands are sufficiently ancient shown the stupendous degradation which they have suffered and their tertiary strata there has also been time for the production endemic species belonging other classes and continents thought that mammals appear and disappear quicker rate than other and lower animals though terrestrial mammals not occur oceanic islands ærial mammals occur almost every island new zealand possesses two bats found nowhere else the world norfolk island the viti archipelago the bonin islands the caroline and marianne archipelagoes and mauritius all possess their peculiar bats why may asked has the supposed creative force produced bats and other mammals remote islands view this question can easily answered for terrestrial mammal can transported across wide space sea but bats can fly across bats have been seen wandering day far over the atlantic ocean and two north american species either regularly occasionally visit bermuda the distance 600 miles from the mainland hear from tomes who has specially studied this family that many the same species have enormous ranges and are found continents and far distant islands hence have only suppose that such wandering species have been modified through natural selection their new homes relation their new position and can understand the presence endemic bats islands with the absence all terrestrial mammals besides the absence terrestrial mammals relation the remoteness islands from continents there also relation certain extent independent distance between the depth the sea separating island from the neighbouring mainland and the presence both the same mammiferous species allied species more less modified condition windsor earl has made some striking observations this head regard the great malay archipelago which traversed near celebes space deep ocean and this space separates two widely distinct mammalian faunas either side the islands are situated moderately deep submarine banks and they are inhabited closely allied identical quadrupeds doubt some few anomalies occur this great archipelago and there much difficulty forming judgment some cases owing the probable naturalisation certain mammals through man agency but shall soon have much light thrown the natural history this archipelago the admirable zeal and researches wallace have not yet had time follow this subject all other quarters the world but far have gone the relation generally holds good see britain separated shallow channel from europe and the mammals are the same both sides meet with analogous facts many islands separated similar channels from australia the west indian islands stand deeply submerged bank nearly 1000 fathoms depth and here find american forms but the species and even the genera are distinct the amount modification all cases depends certain degree the lapse time and during changes level obvious that islands separated shallow channels are more likely have been continuously united within recent period the mainland than islands separated deeper channels can understand the frequent relation between the depth the sea and the degree affinity the mammalian inhabitants islands with those neighbouring continent inexplicable relation the view independent acts creation all the foregoing remarks the inhabitants oceanic islands namely the scarcity kinds the richness endemic forms particular classes sections classes the absence whole groups batrachians and terrestrial mammals notwithstanding the presence ærial bats the singular proportions certain orders plants herbaceous forms having been developed into trees etc seem accord better with the view occasional means transport having been largely efficient the long course time than with the view all our oceanic islands having been formerly connected continuous land with the nearest continent for this latter view the migration would probably have been more complete and modification admitted all the forms life would have been more equally modified accordance with the paramount importance the relation organism organism not deny that there are many and grave difficulties understanding how several the inhabitants the more remote islands whether still retaining the same specific form modified since their arrival could have reached their present homes but the probability many islands having existed halting places which not wreck now remains must not overlooked will here give single instance one the cases difficulty almost all oceanic islands even the most isolated and smallest are inhabited land shells generally endemic species but sometimes species found elsewhere aug gould has given several interesting cases regard the land shells the islands the pacific now notorious that land shells are very easily killed salt their eggs least such have tried sink sea water and are killed yet there must view some unknown but highly efficient means for their transportal would the just hatched young occasionally crawl and adhere the feet birds roosting the ground and thus get transported occurred that land shells when hybernating and having membranous diaphragm over the mouth the shell might floated chinks drifted timber across moderately wide arms the sea and found that several species did this state withstand uninjured immersion sea water during seven days one these shells was the helix pomatia and after had again hybernated put sea water for twenty days and perfectly recovered this species has thick calcareous operculum removed and when had formed new membranous one immersed for fourteen days sea water and recovered and crawled away but more experiments are wanted this head the most striking and important fact for regard the inhabitants islands their affinity those the nearest mainland without being actually the same species numerous instances could given this fact will give only one that the galapagos archipelago situated under the equator between 500 and 600 miles from the shores south america here almost every product the land and water bears the unmistakeable stamp the american continent there are twenty six land birds and twenty five these are ranked gould distinct species supposed have been created here yet the close affinity most these birds american species every character their habits gestures and tones voice was manifest with the other animals and with nearly all the plants shown hooker his admirable memoir the flora this archipelago the naturalist looking the inhabitants these volcanic islands the pacific distant several hundred miles from the continent yet feels that standing american land why should this why should the species which are supposed have been created the galapagos archipelago and nowhere else bear plain stamp affinity those created america there nothing the conditions life the geological nature the islands their height climate the proportions which the several classes are associated together which resembles closely the conditions the south american coast fact there considerable dissimilarity all these respects the other hand there considerable degree resemblance the volcanic nature the soil climate height and size the islands between the galapagos and cape verde archipelagos but what entire and absolute difference their inhabitants the inhabitants the cape verde islands are related those africa like those the galapagos america believe this grand fact can receive sort explanation the ordinary view independent creation whereas the view here maintained obvious that the galapagos islands would likely receive colonists whether occasional means transport formerly continuous land from america and the cape verde islands from africa and that such colonists would liable modification the principle inheritance still betraying their original birthplace many analogous facts could given indeed almost universal rule that the endemic productions islands are related those the nearest continent other near islands the exceptions are few and most them can explained thus the plants kerguelen land though standing nearer africa than america are related and that very closely know from hooker account those america but the view that this island has been mainly stocked seeds brought with earth and stones icebergs drifted the prevailing currents this anomaly disappears new zealand its endemic plants much more closely related australia the nearest mainland than any other region and this what might have been expected but also plainly related south america which although the next nearest continent enormously remote that the fact becomes anomaly but this difficulty almost disappears the view that both new zealand south america and other southern lands were long ago partially stocked from nearly intermediate though distant point namely from the antarctic islands when they were clothed with vegetation before the commencement the glacial period the affinity which though feeble assured hooker real between the flora the south western corner australia and the cape good hope far more remarkable case and present inexplicable but this affinity confined the plants and will not doubt some day explained the law which causes the inhabitants archipelago though specifically distinct closely allied those the nearest continent sometimes see displayed small scale yet most interesting manner within the limits the same archipelago thus the several islands the galapagos archipelago are tenanted have elsewhere shown quite marvellous manner very closely related species that the inhabitants each separate island though mostly distinct are related incomparably closer degree each other than the inhabitants any other part the world and this just what might have been expected view for the islands are situated near each other that they would almost certainly receive immigrants from the same original source from each other but this dissimilarity between the endemic inhabitants the islands may used argument against views for may asked how has happened the several islands situated within sight each other having the same geological nature the same height climate etc that many the immigrants should have been differently modified though only small degree this long appeared great difficulty but arises chief part from the deeply seated error considering the physical conditions country the most important for its inhabitants whereas cannot think disputed that the nature the other inhabitants with which each has compete least important and generally far more important element success now look those inhabitants the galapagos archipelago which are found other parts the world laying one side for the moment the endemic species which cannot here fairly included are considering how they have come modified since their arrival find considerable amount difference the several islands this difference might indeed have been expected the view the islands having been stocked occasional means transport seed for instance one plant having been brought one island and that another plant another island hence when former times immigrant settled any one more the islands when subsequently spread from one island another would undoubtedly exposed different conditions life the different islands for would have compete with different sets organisms plant for instance would find the best fitted ground more perfectly occupied distinct plants one island than another and would exposed the attacks somewhat different enemies then varied natural selection would probably favour different varieties the different islands some species however might spread and yet retain the same character throughout the group just see continents some species spreading widely and remaining the same the really surprising fact this case the galapagos archipelago and lesser degree some analogous instances that the new species formed the separate islands have not quickly spread the other islands but the islands though sight each other are separated deep arms the sea most cases wider than the british channel and there reason suppose that they have any former period been continuously united the currents the sea are rapid and sweep across the archipelago and gales wind are extraordinarily rare that the islands are far more effectually separated from each other than they appear map nevertheless good many species both those found other parts the world and those confined the archipelago are common the several islands and may infer from certain facts that these have probably spread from some one island the others but often take think erroneous view the probability closely allied species invading each other territory when put into free intercommunication undoubtedly one species has any advantage whatever over another will very brief time wholly part supplant but both are equally well fitted for their own places nature both probably will hold their own places and keep separate for almost any length time being familiar with the fact that many species naturalised through man agency have spread with astonishing rapidity over new countries are apt infer that most species would thus spread but should remember that the forms which become naturalised new countries are not generally closely allied the aboriginal inhabitants but are very distinct species belonging large proportion cases shown alph candolle distinct genera the galapagos archipelago many even the birds though well adapted for flying from island island are distinct each thus there are three closely allied species mocking thrush each confined its own island now let suppose the mocking thrush chatham island blown charles island which has its own mocking thrush why should succeed establishing itself there may safely infer that charles island well stocked with its own species for annually more eggs are laid there than can possibly reared and may infer that the mocking thrush peculiar charles island least well fitted for its home the species peculiar chatham island sir lyell and wollaston have communicated remarkable fact bearing this subject namely that madeira and the adjoining islet porto santo possess many distinct but representative land shells some which live crevices stone and although large quantities stone are annually transported from porto santo madeira yet this latter island has not become colonised the porto santo species nevertheless both islands have been colonised some european land shells which doubt had some advantage over the indigenous species from these considerations think need not greatly marvel the endemic and representative species which inhabit the several islands the galapagos archipelago not having universally spread from island island many other instances the several districts the same continent pre occupation has probably played important part checking the commingling species under the same conditions life thus the south east and south west corners australia have nearly the same physical conditions and are united continuous land yet they are inhabited vast number distinct mammals birds and plants the principle which determines the general character the fauna and flora oceanic islands namely that the inhabitants when not identically the same yet are plainly related the inhabitants that region whence colonists could most readily have been derived the colonists having been subsequently modified and better fitted their new homes the widest application throughout nature see this every mountain every lake and marsh for alpine species excepting far the same forms chiefly plants have spread widely throughout the world during the recent glacial epoch are related those the surrounding lowlands thus have south america alpine humming birds alpine rodents alpine plants etc all strictly american forms and obvious that mountain became slowly upheaved would naturally colonised from the surrounding lowlands with the inhabitants lakes and marshes excepting far great facility transport has given the same general forms the whole world see this same principle the blind animals inhabiting the caves america and europe other analogous facts could given and will believe universally found true that wherever two regions let them ever distant many closely allied representative species occur there will likewise found some identical species showing accordance with the foregoing view that some former period there has been intercommunication migration between the two regions and wherever many closely allied species occur there will found many forms which some naturalists rank distinct species and some varieties these doubtful forms showing the steps the process modification this relation between the power and extent migration species either the present time some former period under different physical conditions and the existence remote points the world other species allied shown another and more general way gould remarked long ago that those genera birds which range over the world many the species have very wide ranges can hardly doubt that this rule generally true though would difficult prove amongst mammals see strikingly displayed bats and lesser degree the felidæ and canidæ see compare the distribution butterflies and beetles with most fresh water productions which many genera range over the world and many individual species have enormous ranges not meant that world ranging genera all the species have wide range even that they have average wide range but only that some the species range very widely for the facility with which widely ranging species vary and give rise new forms will largely determine their average range for instance two varieties the same species inhabit america and europe and the species thus has immense range but the variation had been little greater the two varieties would have been ranked distinct species and the common range would have been greatly reduced still less meant that species which apparently has the capacity crossing barriers and ranging widely the case certain powerfully winged birds will necessarily range widely for should never forget that range widely implies not only the power crossing barriers but the more important power being victorious distant lands the struggle for life with foreign associates but the view all the species genus having descended from single parent though now distributed the most remote points the world ought find and believe general rule find that some least the species range very widely for necessary that the unmodified parent should range widely undergoing modification during its diffusion and should place itself under diverse conditions favourable for the conversion its offspring firstly into new varieties and ultimately into new species considering the wide distribution certain genera should bear mind that some are extremely ancient and must have branched off from common parent remote epoch that such cases there will have been ample time for great climatal and geographical changes and for accidents transport and consequently for the migration some the species into all quarters the world where they may have become slightly modified relation their new conditions there also some reason believe from geological evidence that organisms low the scale within each great class generally change slower rate than the higher forms and consequently the lower forms will have had better chance ranging widely and still retaining the same specific character this fact together with the seeds and eggs many low forms being very minute and better fitted for distant transportation probably accounts for law which has long been observed and which has lately been admirably discussed alph candolle regard plants namely that the lower any group organisms the more widely apt range the relations just discussed namely low and slowly changing organisms ranging more widely than the high some the species widely ranging genera themselves ranging widely such facts alpine lacustrine and marsh productions being related with the exceptions before specified those the surrounding low lands and dry lands though these stations are different the very close relation the distinct species which inhabit the islets the same archipelago and especially the striking relation the inhabitants each whole archipelago island those the nearest mainland are think utterly inexplicable the ordinary view the independent creation each species but are explicable the view colonisation from the nearest and readiest source together with the subsequent modification and better adaptation the colonists their new homes summary last and present chapters these chapters have endeavoured show that make due allowance for our ignorance the full effects all the changes climate and the level the land which have certainly occurred within the recent period and other similar changes which may have occurred within the same period remember how profoundly ignorant are with respect the many and curious means occasional transport subject which has hardly ever been properly experimentised bear mind how often species may have ranged continuously over wide area and then have become extinct the intermediate tracts think the difficulties believing that all the individuals the same species wherever located have descended from the same parents are not insuperable and are led this conclusion which has been arrived many naturalists under the designation single centres creation some general considerations more especially from the importance barriers and from the analogical distribution sub genera genera and families with respect the distinct species the same genus which theory must have spread from one parent source make the same allowances before for our ignorance and remember that some forms life change most slowly enormous periods time being thus granted for their migration not think that the difficulties are insuperable though they often are this case and that the individuals the same species extremely grave exemplifying the effects climatal changes distribution have attempted show how important has been the influence the modern glacial period which fully convinced simultaneously affected the whole world least great meridional belts showing how diversified are the means occasional transport have discussed some little length the means dispersal fresh water productions the difficulties not insuperable admitting that the long course time the individuals the same species and likewise allied species have proceeded from some one source then think all the grand leading facts geographical distribution are explicable the theory migration generally the more dominant forms life together with subsequent modification and the multiplication new forms can thus understand the high importance barriers whether land water which separate our several zoological and botanical provinces can thus understand the localisation sub genera genera and families and how that under different latitudes for instance south america the inhabitants the plains and mountains the forests marshes and deserts are mysterious manner linked together affinity and are likewise linked the extinct beings which formerly inhabited the same continent bearing mind that the mutual relations organism organism are the highest importance can see why two areas having nearly the same physical conditions should often inhabited very different forms life for according the length time which has elapsed since new inhabitants entered one region according the nature the communication which allowed certain forms and not others enter either greater lesser numbers according not those which entered happened come more less direct competition with each other and with the aborigines and according the immigrants were capable varying more less rapidly there would ensue different regions independently their physical conditions infinitely diversified conditions life there would almost endless amount organic action and reaction and should find find some groups beings greatly and some only slightly modified some developed great force some existing scanty numbers the different great geographical provinces the world these same principles can understand have endeavoured show why oceanic islands should have few inhabitants but these great number should endemic peculiar and why relation the means migration one group beings even within the same class should have all its species endemic and another group should have all its species common other quarters the world can see why whole groups organisms batrachians and terrestrial mammals should absent from oceanic islands whilst the most isolated islands possess their own peculiar species ærial mammals bats can see why there should some relation between the presence mammals more less modified condition and the depth the sea between island and the mainland can clearly see why all the inhabitants archipelago though specifically distinct the several islets should closely related each other and likewise related but less closely those the nearest continent other source whence immigrants were probably derived can see why two areas however distant from each other there should correlation the presence identical species varieties doubtful species and distinct but representative species the late edward forbes often insisted there striking parallelism the laws life throughout time and space the laws governing the succession forms past times being nearly the same with those governing the present time the differences different areas see this many facts the endurance each species and group species continuous time for the exceptions the rule are few that they may fairly attributed our not having yet discovered intermediate deposit the forms which are therein absent but which occur above and below space certainly the general rule that the area inhabited single species group species continuous and the exceptions which are not rare may have attempted show accounted for migration some former period under different conditions occasional means transport and the species having become extinct the intermediate tracts both time and space species and groups species have their points maximum development groups species belonging either certain period time certain area are often characterised trifling characters common sculpture colour looking the long succession ages now looking distant provinces throughout the world find that some organisms differ little whilst others belonging different class different order even only different family the same order differ greatly both time and space the lower members each class generally change less than the higher but there are both cases marked exceptions the rule theory these several relations throughout time and space are intelligible for whether look the forms life which have changed during successive ages within the same quarter the world those which have changed after having migrated into distant quarters both cases the forms within each class have been connected the same bond ordinary generation and the more nearly any two forms are related blood the nearer they will generally stand each other time and space both cases the laws variation have been the same and modifications have been accumulated the same power natural selection chapter xiii mutual affinities organic beings morphology embryology rudimentary organs classification groups subordinate groups natural system rules and difficulties classification explained the theory descent with modification classification varieties descent always used classification analogical adaptive characters affinities general complex and radiating extinction separates and defines groups morphology between members the same class between parts the same individual embryology laws explained variations not supervening early age and being inherited corresponding age rudimentary organs their origin explained summary from the first dawn life all organic beings are found resemble each other descending degrees that they can classed groups under groups this classification evidently not arbitrary like the grouping the stars constellations the existence groups would have been simple signification one group had been exclusively fitted inhabit the land and another the water one feed flesh another vegetable matter and but the case widely different nature for notorious how commonly members even the same subgroup have different habits our second and fourth chapters variation and natural selection have attempted show that the widely ranging the much diffused and common that the dominant species belonging the larger genera which vary most the varieties incipient species thus produced ultimately become converted believe into new and distinct species and these the principle inheritance tend produce other new and dominant species consequently the groups which are now large and which generally include many dominant species tend increasing indefinitely size further attempted show that from the varying descendants each species trying occupy many and different places possible the economy nature there constant tendency their characters diverge this conclusion was supported looking the great diversity the forms life which any small area come into the closest competition and looking certain facts naturalisation attempted also show that there constant tendency the forms which are increasing number and diverging character supplant and exterminate the less divergent the less improved and preceding forms request the reader turn the diagram illustrating the action formerly explained these several principles and will see that the inevitable result that the modified descendants proceeding from one progenitor become broken into groups subordinate groups the diagram each letter the uppermost line may represent genus including several species and all the genera this line form together one class for all have descended from one ancient but unseen parent and consequently have inherited something common but the three genera the left hand have this same principle much common and form sub family distinct from that including the next two genera the right hand which diverged from common parent the fifth stage descent these five genera have also much though less common and they form family distinct from that including the three genera still further the right hand which diverged still earlier period and all these genera descended from form order distinct from the genera descended from that here have many species descended from single progenitor grouped into genera and the genera are included subordinate sub families families and orders all united into one class thus the grand fact natural history the subordination group under group which from its familiarity does not always sufficiently strike judgment fully explained naturalists try arrange the species genera and families each class what called the natural system but what meant this system some authors look merely scheme for arranging together those living objects which are most alike and for separating those which are most unlike artificial means for enunciating briefly possible general propositions that one sentence give the characters common for instance all mammals another those common all carnivora another those common the dog genus and then adding single sentence full description given each kind dog the ingenuity and utility this system are indisputable but many naturalists think that something more meant the natural system they believe that reveals the plan the creator but unless specified whether order time space what else meant the plan the creator seems that nothing thus added our knowledge such expressions that famous one linnæus and which often meet with more less concealed form that the characters not make the genus but that the genus gives the characters seem imply that something more included our classification than mere resemblance believe that something more included and that propinquity descent the only known cause the similarity organic beings the bond hidden various degrees modification which partially revealed our classifications let now consider the rules followed classification and the difficulties which are encountered the view that classification either gives some unknown plan creation simply scheme for enunciating general propositions and placing together the forms most like each other might have been thought and was ancient times thought that those parts the structure which determined the habits life and the general place each being the economy nature would very high importance classification nothing can more false one regards the external similarity mouse shrew dugong whale whale fish any importance these resemblances though intimately connected with the whole life the being are ranked merely adaptive analogical characters but the consideration these resemblances shall have recur may even given general rule that the less any part the organisation concerned with special habits the more important becomes for classification instance owen speaking the dugong says the generative organs being those which are most remotely related the habits and food animal have always regarded affording very clear indications its true affinities are least likely the modifications these organs mistake merely adaptive for essential character with plants how remarkable that the organs vegetation which their whole life depends are little signification excepting the first main divisions whereas the organs reproduction with their product the seed are paramount importance must not therefore classifying trust resemblances parts the organisation however important they may for the welfare the being relation the outer world perhaps from this cause has partly arisen that almost all naturalists lay the greatest stress resemblances organs high vital physiological importance doubt this view the classificatory importance organs which are important generally but means always true but their importance for classification believe depends their greater constancy throughout large groups species and this constancy depends such organs having generally been subjected less change the adaptation the species their conditions life that the mere physiological importance organ does not determine its classificatory value almost shown the one fact that allied groups which the same organ have every reason suppose has nearly the same physiological value its classificatory value widely different naturalist can have worked any group without being struck with this fact and has been most fully acknowledged the writings almost every author will suffice quote the highest authority robert brown who speaking certain organs the proteaceæ says their generic importance like that all their parts not only this but apprehend every natural family very unequal and some cases seems entirely lost again another work says the genera the connaraceæ differ having one more ovaria the existence absence albumen the imbricate valvular æstivation any one these characters singly frequently more than generic importance though here even when all taken together they appear insufficient separate cnestis from connarus give example amongst insects one great division the hymenoptera the antennæ westwood has remarked are most constant structure another division they differ much and the differences are quite subordinate value classification yet one probably will say that the antennæ these two divisions the same order are unequal physiological importance any number instances could given the varying importance for classification the same important organ within the same group beings again one will say that rudimentary atrophied organs are high physiological vital importance yet undoubtedly organs this condition are often high value classification one will dispute that the rudimentary teeth the upper jaws young ruminants and certain rudimentary bones the leg are highly serviceable exhibiting the close affinity between ruminants and pachyderms robert brown has strongly insisted the fact that the rudimentary florets are the highest importance the classification the grasses numerous instances could given characters derived from parts which must considered very trifling physiological importance but which are universally admitted highly serviceable the definition whole groups for instance whether not there open passage from the nostrils the mouth the only character according owen which absolutely distinguishes fishes and reptiles the inflection the angle the jaws marsupials the manner which the wings insects are folded mere colour certain algæ mere pubescence parts the flower grasses the nature the dermal covering hair feathers the vertebrata the ornithorhynchus had been covered with feathers instead hair this external and trifling character would think have been considered naturalists important aid determining the degree affinity this strange creature birds and reptiles approach structure any one internal and important organ the importance for classification trifling characters mainly depends their being correlated with several other characters more less importance the value indeed aggregate characters very evident natural history hence has often been remarked species may depart from its allies several characters both high physiological importance and almost universal prevalence and yet leave doubt where should ranked hence also has been found that classification founded any single character however important that may has always failed for part the organisation universally constant the importance aggregate characters even when none are important alone explains think that saying linnæus that the characters not give the genus but the genus gives the characters for this saying seems founded appreciation many trifling points resemblance too slight defined certain plants belonging the malpighiaceæ bear perfect and degraded flowers the latter jussieu has remarked the greater number the characters proper the species the genus the family the class disappear and thus laugh our classification but when aspicarpa produced france during several years only degraded flowers departing wonderfully number the most important points structure from the proper type the order yet richard sagaciously saw jussieu observes that this genus should still retained amongst the malpighiaceæ this case seems well illustrate the spirit with which our classifications are sometimes necessarily founded practically when naturalists are work they not trouble themselves about the physiological value the characters which they use defining group allocating any particular species they find character nearly uniform and common great number forms and not common others they use one high value common some lesser number they use subordinate value this principle has been broadly confessed some naturalists the true one and none more clearly than that excellent botanist aug hilaire certain characters are always found correlated with others though apparent bond connexion can discovered between them especial value set them most groups animals important organs such those for propelling the blood for ærating those for propagating the race are found nearly uniform they are considered highly serviceable classification but some groups animals all these the most important vital organs are found offer characters quite subordinate value can see why characters derived from the embryo should equal importance with those derived from the adult for our classifications course include all ages each species but means obvious the ordinary view why the structure the embryo should more important for this purpose than that the adult which alone plays its full part the economy nature yet has been strongly urged those great naturalists milne edwards and agassiz that embryonic characters are the most important any the classification animals and this doctrine has very generally been admitted true the same fact holds good with flowering plants which the two main divisions have been founded characters derived from the embryo the number and position the embryonic leaves cotyledons and the mode development the plumule and radicle our discussion embryology shall see why such characters are valuable the view classification tacitly including the idea descent our classifications are often plainly influenced chains affinities nothing can easier than define number characters common all birds but the case crustaceans such definition has hitherto been found impossible there are crustaceans the opposite ends the series which have hardly character common yet the species both ends from being plainly allied others and these others and onwards can recognised unequivocally belonging this and other class the articulata geographical distribution has often been used though perhaps not quite logically classification more especially very large groups closely allied forms temminck insists the utility even necessity this practice certain groups birds and has been followed several entomologists and botanists finally with respect the comparative value the various groups species such orders sub orders families sub families and genera they seem least present almost arbitrary several the best botanists such bentham and others have strongly insisted their arbitrary value instances could given amongst plants and insects group forms first ranked practised naturalists only genus and then raised the rank sub family family and this has been done not because further research has detected important structural differences first overlooked but because numerous allied species with slightly different grades difference have been subsequently discovered all the foregoing rules and aids and difficulties classification are explained not greatly deceive myself the view that the natural system founded descent with modification that the characters which naturalists consider showing true affinity between any two more species are those which have been inherited from common parent and far all true classification genealogical that community descent the hidden bond which naturalists have been unconsciously seeking and not some unknown plan creation the enunciation general propositions and the mere putting together and separating objects more less alike but must explain meaning more fully believe that the arrangement the groups within each class due subordination and relation the other groups must strictly genealogical order natural but that the amount difference the several branches groups though allied the same degree blood their common progenitor may differ greatly being due the different degrees modification which they have undergone and this expressed the forms being ranked under different genera families sections orders the reader will best understand what meant will take the trouble referring the diagram the fourth chapter will suppose the letters represent allied genera which lived during the silurian epoch and these have descended from species which existed unknown anterior period species three these genera and have transmitted modified descendants the present day represented the fifteen genera the uppermost horizontal line now all these modified descendants from single species are represented related blood descent the same degree they may metaphorically called cousins the same millionth degree yet they differ widely and different degrees from each other the forms descended from now broken into two three families constitute distinct order from those descended from also broken into two families nor can the existing species descended from ranked the same genus with the parent those from with the parent but the existing genus f14 may supposed have been but slightly modified and will then rank with the parent genus just some few still living organic beings belong silurian genera that the amount value the differences between organic beings all related each other the same degree blood has come widely different nevertheless their genealogical arrangement remains strictly true not only the present time but each successive period descent all the modified descendants from will have inherited something common from their common parent will all the descendants from will with each subordinate branch descendants each successive period however choose suppose that any the descendants have been much modified have more less completely lost traces their parentage this case their places natural classification will have been more less completely lost sometimes seems have occurred with existing organisms all the descendants the genus along its whole line descent are supposed have been but little modified and they yet form single genus but this genus though much isolated will still occupy its proper intermediate position for originally was intermediate character between and and the several genera descended from these two genera will have inherited certain extent their characters this natural arrangement shown far possible paper the diagram but much too simple manner branching diagram had not been used and only the names the groups had been written linear series would have been still less possible have given natural arrangement and notoriously not possible represent series flat surface the affinities which discover nature amongst the beings the same group thus the view which hold the natural system genealogical its arrangement like pedigree but the degrees modification which the different groups have undergone have expressed ranking them under different called genera sub families families sections orders and classes may worth while illustrate this view classification taking the case languages possessed perfect pedigree mankind genealogical arrangement the races man would afford the best classification the various languages now spoken throughout the world and all extinct languages and all intermediate and slowly changing dialects had included such arrangement would think the only possible one yet might that some very ancient language had altered little and had given rise few new languages whilst others owing the spreading and subsequent isolation and states civilisation the several races descended from common race had altered much and had given rise many new languages and dialects the various degrees difference the languages from the same stock would have expressed groups subordinate groups but the proper even only possible arrangement would still genealogical and this would strictly natural would connect together all languages extinct and modern the closest affinities and would give the filiation and origin each tongue confirmation this view let glance the classification varieties which are believed known have descended from one species these are grouped under species with sub varieties under varieties and with our domestic productions several other grades difference are requisite have seen with pigeons the origin the existence groups subordinate groups the same with varieties with species namely closeness descent with various degrees modification nearly the same rules are followed classifying varieties with species authors have insisted the necessity classing varieties natural instead artificial system are cautioned for instance not class two varieties the pine apple together merely because their fruit though the most important part happens nearly identical one puts the swedish and common turnips together though the esculent and thickened stems are similar whatever part found most constant used classing varieties thus the great agriculturist marshall says the horns are very useful for this purpose with cattle because they are less variable than the shape colour the body etc whereas with sheep the horns are much less serviceable because less constant classing varieties apprehend had real pedigree genealogical classification would universally preferred and has been attempted some authors for might feel sure whether there had been more less modification the principle inheritance would keep the forms together which were allied the greatest number points tumbler pigeons though some sub varieties differ from the others the important character having longer beak yet all are kept together from having the common habit tumbling but the short faced breed has nearly quite lost this habit nevertheless without any reasoning thinking the subject these tumblers are kept the same group because allied blood and alike some other respects could proved that the hottentot had descended from the negro think would classed under the negro group however much might differ colour and other important characters from negroes with species state nature every naturalist has fact brought descent into his classification for includes his lowest grade that species the two sexes and how enormously these sometimes differ the most important characters known every naturalist scarcely single fact can predicated common the males and hermaphrodites certain cirripedes when adult and yet one dreams separating them the naturalist includes one species the several larval stages the same individual however much they may differ from each other and from the adult likewise includes the called alternate generations steenstrup which can only technical sense considered the same individual includes monsters includes varieties not solely because they closely resemble the parent form but because they are descended from who believes that the cowslip descended from the primrose conversely ranks them together single species and gives single definition soon three orchidean forms monochanthus myanthus and catasetum which had previously been ranked three distinct genera were known sometimes produced the same spike they were immediately included single species but may asked what ought could proved that one species kangaroo had been produced long course modification from bear ought rank this one species with bears and what should with the other species the supposition course preposterous and might answer the argumentum hominem and ask what should done perfect kangaroo were seen come out the womb bear according all analogy would ranked with bears but then assuredly all the other species the kangaroo family would have classed under the bear genus the whole case preposterous for where there has been close descent common there will certainly close resemblance affinity descent has universally been used classing together the individuals the same species though the males and females and larvæ are sometimes extremely different and has been used classing varieties which have undergone certain and sometimes considerable amount modification may not this same element descent have been unconsciously used grouping species under genera and genera under higher groups though these cases the modification has been greater degree and has taken longer time complete believe has thus been unconsciously used and only thus can understand the several rules and guides which have been followed our best systematists have written pedigrees have make out community descent resemblances any kind therefore choose those characters which far can judge are the least likely have been modified relation the conditions life which each species has been recently exposed rudimentary structures this view are good even sometimes better than other parts the organisation care not how trifling character may let the mere inflection the angle the jaw the manner which insect wing folded whether the skin covered hair feathers prevail throughout many and different species especially those having very different habits life assumes high value for can account for its presence many forms with such different habits only its inheritance from common parent may err this respect regard single points structure but when several characters let them ever trifling occur together throughout large group beings having different habits may feel almost sure the theory descent that these characters have been inherited from common ancestor and know that such correlated aggregated characters have especial value classification can understand why species group species may depart several its most important characteristics from its allies and yet safely classed with them this may safely done and often done long sufficient number characters let them ever unimportant betrays the hidden bond community descent let two forms have not single character common yet these extreme forms are connected together chain intermediate groups may once infer their community descent and put them all into the same class find organs high physiological importance those which serve preserve life under the most diverse conditions existence are generally the most constant attach especial value them but these same organs another group section group are found differ much once value them less our classification shall hereafter think clearly see why embryological characters are such high classificatory importance geographical distribution may sometimes brought usefully into play classing large and widely distributed genera because all the species the same genus inhabiting any distinct and isolated region have all probability descended from the same parents can understand these views the very important distinction between real affinities and analogical adaptive resemblances lamarck first called attention this distinction and has been ably followed macleay and others the resemblance the shape the body and the fin like anterior limbs between the dugong which pachydermatous animal and the whale and between both these mammals and fishes analogical amongst insects there are innumerable instances thus linnæus misled external appearances actually classed homopterous insect moth see something the same kind even our domestic varieties the thickened stems the common and swedish turnip the resemblance the greyhound and racehorse hardly more fanciful than the analogies which have been drawn some authors between very distinct animals view characters being real importance for classification only far they reveal descent can clearly understand why analogical adaptive character although the utmost importance the welfare the being are almost valueless the systematist for animals belonging two most distinct lines descent may readily become adapted similar conditions and thus assume close external resemblance but such resemblances will not reveal will rather tend conceal their blood relationship their proper lines descent can also understand the apparent paradox that the very same characters are analogical when one class order compared with another but give true affinities when the members the same class order are compared one with another thus the shape the body and fin like limbs are only analogical when whales are compared with fishes being adaptations both classes for swimming through the water but the shape the body and fin like limbs serve characters exhibiting true affinity between the several members the whale family for these cetaceans agree many characters great and small that cannot doubt that they have inherited their general shape body and structure limbs from common ancestor with fishes members distinct classes have often been adapted successive slight modifications live under nearly similar circumstances inhabit for instance the three elements land air and water can perhaps understand how that numerical parallelism has sometimes been observed between the sub groups distinct classes naturalist struck parallelism this nature any one class arbitrarily raising sinking the value the groups other classes and all our experience shows that this valuation has hitherto been arbitrary could easily extend the parallelism over wide range and thus the septenary quinary quaternary and ternary classifications have probably arisen the modified descendants dominant species belonging the larger genera tend inherit the advantages which made the groups which they belong large and their parents dominant they are almost sure spread widely and seize more and more places the economy nature the larger and more dominant groups thus tend increasing size and they consequently supplant many smaller and feebler groups thus can account for the fact that all organisms recent and extinct are included under few great orders under still fewer classes and all one great natural system showing how few the higher groups are number and how widely spread they are throughout the world the fact striking that the discovery australia has not added single insect belonging new order and that the vegetable kingdom learn from hooker has added only two three orders small size the chapter geological succession attempted show the principle each group having generally diverged much character during the long continued process modification how that the more ancient forms life often present characters some slight degree intermediate between existing groups few old and intermediate parent forms having occasionally transmitted the present day descendants but little modified will give our called osculant aberrant groups the more aberrant any form the greater must the number connecting forms which theory have been exterminated and utterly lost and have some evidence aberrant forms having suffered severely from extinction for they are generally represented extremely few species and such species occur are generally very distinct from each other which again implies extinction the genera ornithorhynchus and lepidosiren for example would not have been less aberrant had each been represented dozen species instead single one but such richness species find after some investigation does not commonly fall the lot aberrant genera can think account for this fact only looking aberrant forms failing groups conquered more successful competitors with few members preserved some unusual coincidence favourable circumstances waterhouse has remarked that when member belonging one group animals exhibits affinity quite distinct group this affinity most cases general and not special thus according waterhouse all rodents the bizcacha most nearly related marsupials but the points which approaches this order its relations are general and not any one marsupial species more than another the points affinity the bizcacha marsupials are believed real and not merely adaptive they are due theory inheritance common therefore must suppose either that all rodents including the bizcacha branched off from some very ancient marsupial which will have had character some degree intermediate with respect all existing marsupials that both rodents and marsupials branched off from common progenitor and that both groups have since undergone much modification divergent directions either view may suppose that the bizcacha has retained inheritance more the character its ancient progenitor than have other rodents and therefore will not specially related any one existing marsupial but indirectly all nearly all marsupials from having partially retained the character their common progenitor early member the group the other hand all marsupials waterhouse has remarked the phascolomys resembles most nearly not any one species but the general order rodents this case however may strongly suspected that the resemblance only analogical owing the phascolomys having become adapted habits like those rodent the elder candolle has made nearly similar observations the general nature the affinities distinct orders plants the principle the multiplication and gradual divergence character the species descended from common parent together with their retention inheritance some characters common can understand the excessively complex and radiating affinities which all the members the same family higher group are connected together for the common parent whole family species now broken extinction into distinct groups and sub groups will have transmitted some its characters modified various ways and degrees all and the several species will consequently related each other circuitous lines affinity various lengths may seen the diagram often referred mounting through many predecessors difficult show the blood relationship between the numerous kindred any ancient and noble family even the aid genealogical tree and almost impossible this without this aid can understand the extraordinary difficulty which naturalists have experienced describing without the aid diagram the various affinities which they perceive between the many living and extinct members the same great natural class extinction have seen the fourth chapter has played important part defining and widening the intervals between the several groups each class may thus account even for the distinctness whole classes from each other for instance birds from all other vertebrate animals the belief that many ancient forms life have been utterly lost through which the early progenitors birds were formerly connected with the early progenitors the other vertebrate classes there has been less entire extinction the forms life which once connected fishes with batrachians there has been still less some other classes that the crustacea for here the most wonderfully diverse forms are still tied together long but broken chain affinities extinction has only separated groups has means made them for every form which has ever lived this earth were suddenly reappear though would quite impossible give definitions which each group could distinguished from other groups all would blend together steps fine those between the finest existing varieties nevertheless natural classification least natural arrangement would possible shall see this turning the diagram the letters may represent eleven silurian genera some which have produced large groups modified descendants every intermediate link between these eleven genera and their primordial parent and every intermediate link each branch and sub branch their descendants may supposed still alive and the links fine those between the finest varieties this case would quite impossible give any definition which the several members the several groups could distinguished from their more immediate parents these parents from their ancient and unknown progenitor yet the natural arrangement the diagram would still hold good and the principle inheritance all the forms descended from from would have something common tree can specify this that branch though the actual fork the two unite and blend together could not have said define the several groups but could pick out types forms representing most the characters each group whether large small and thus give general idea the value the differences between them this what should driven were ever succeed collecting all the forms any class which have lived throughout all time and space shall certainly never succeed making perfect collection nevertheless certain classes are tending this direction and milne edwards has lately insisted able paper the high importance looking types whether not can separate and define the groups which such types belong finally have seen that natural selection which results from the struggle for existence and which almost inevitably induces extinction and divergence character the many descendants from one dominant parent species explains that great and universal feature the affinities all organic beings namely their subordination group under group use the element descent classing the individuals both sexes and all ages although having few characters common under one species use descent classing acknowledged varieties however different they may from their parent and believe this element descent the hidden bond connexion which naturalists have sought under the term the natural system this idea the natural system being far has been perfected genealogical its arrangement with the grades difference between the descendants from common parent expressed the terms genera families orders etc can understand the rules which are compelled follow our classification can understand why value certain resemblances far more than others why are permitted use rudimentary and useless organs others trifling physiological importance why comparing one group with distinct group summarily reject analogical adaptive characters and yet use these same characters within the limits the same group can clearly see how that all living and extinct forms can grouped together one great system and how the several members each class are connected together the most complex and radiating lines affinities shall never probably disentangle the inextricable web affinities between the members any one class but when have distinct object view and not look some unknown plan creation may hope make sure but slow progress morphology have seen that the members the same class independently their habits life resemble each other the general plan their organisation this resemblance often expressed the term unity type saying that the several parts and organs the different species the class are homologous the whole subject included under the general name morphology this the most interesting department natural history and may said its very soul what can more curious than that the hand man formed for grasping that mole for digging the leg the horse the paddle the porpoise and the wing the bat should all constructed the same pattern and should include the same bones the same relative positions geoffroy hilaire has insisted strongly the high importance relative connexion homologous organs the parts may change almost any extent form and size and yet they always remain connected together the same order never find for instance the bones the arm and forearm the thigh and leg transposed hence the same names can given the homologous bones widely different animals see the same great law the construction the mouths insects what can more different than the immensely long spiral proboscis sphinx moth the curious folded one bee bug and the great jaws beetle yet all these organs serving for such different purposes are formed infinitely numerous modifications upper lip mandibles and two pairs maxillæ analogous laws govern the construction the mouths and limbs crustaceans with the flowers plants nothing can more hopeless than attempt explain this similarity pattern members the same class utility the doctrine final causes the hopelessness the attempt has been expressly admitted owen his most interesting work the nature limbs the ordinary view the independent creation each being can only say that that has pleased the creator construct each animal and plant the explanation manifest the theory the natural selection successive slight modifications each modification being profitable some way the modified form but often affecting correlation growth other parts the organisation changes this nature there will little tendency modify the original pattern transpose parts the bones limb might shortened and widened any extent and become gradually enveloped thick membrane serve fin webbed foot might have all its bones certain bones lengthened any extent and the membrane connecting them increased any extent serve wing yet all this great amount modification there will tendency alter the framework bones the relative connexion the several parts suppose that the ancient progenitor the archetype may called all mammals had its limbs constructed the existing general pattern for whatever purpose they served can once perceive the plain signification the homologous construction the limbs throughout the whole class with the mouths insects have only suppose that their common progenitor had upper lip mandibles and two pair maxillæ these parts being perhaps very simple form and then natural selection will account for the infinite diversity structure and function the mouths insects nevertheless conceivable that the general pattern organ might become much obscured finally lost the atrophy and ultimately the complete abortion certain parts the soldering together other parts and the doubling multiplication others variations which know within the limits possibility the paddles the extinct gigantic sea lizards and the mouths certain suctorial crustaceans the general pattern seems have been thus certain extent obscured there another and equally curious branch the present subject namely the comparison not the same part different members class but the different parts organs the same individual most physiologists believe that the bones the skull are homologous with that correspond number and relative connexion with the elemental parts certain number vertebræ the anterior and posterior limbs each member the vertebrate and articulate classes are plainly homologous see the same law comparing the wonderfully complex jaws and legs crustaceans familiar almost every one that flower the relative position the sepals petals stamens and pistils well their intimate structure are intelligible the view that they consist metamorphosed leaves arranged spire monstrous plants often get direct evidence the possibility one organ being transformed into another and can actually see embryonic crustaceans and many other animals and flowers that organs which when mature become extremely different are early stage growth exactly alike how inexplicable are these facts the ordinary view creation why should the brain enclosed box composed such numerous and such extraordinarily shaped pieces bone owen has remarked the benefit derived from the yielding the separate pieces the act parturition mammals will means explain the same construction the skulls birds why should similar bones have been created the formation the wing and leg bat used they are for such totally different purposes why should one crustacean which has extremely complex mouth formed many parts consequently always have fewer legs conversely those with many legs have simpler mouths why should the sepals petals stamens and pistils any individual flower though fitted for such widely different purposes all constructed the same pattern the theory natural selection can satisfactorily answer these questions the vertebrata see series internal vertebræ bearing certain processes and appendages the articulata see the body divided into series segments bearing external appendages and flowering plants see series successive spiral whorls leaves indefinite repetition the same part organ the common characteristic owen has observed all low little modified forms therefore may readily believe that the unknown progenitor the vertebrata possessed many vertebræ the unknown progenitor the articulata many segments and the unknown progenitor flowering plants many spiral whorls leaves have formerly seen that parts many times repeated are eminently liable vary number and structure consequently quite probable that natural selection during long continued course modification should have seized certain number the primordially similar elements many times repeated and have adapted them the most diverse purposes and the whole amount modification will have been effected slight successive steps need not wonder discovering such parts organs certain degree fundamental resemblance retained the strong principle inheritance the great class molluscs though can homologise the parts one species with those another and distinct species can indicate but few serial homologies that are seldom enabled say that one part organ homologous with another the same individual and can understand this fact for molluscs even the lowest members the class not find nearly much indefinite repetition any one part find the other great classes the animal and vegetable kingdoms naturalists frequently speak the skull formed metamorphosed vertebræ the jaws crabs metamorphosed legs the stamens and pistils flowers metamorphosed leaves but would these cases probably more correct professor huxley has remarked speak both skull and vertebræ both jaws and legs etc having been metamorphosed not one from the other but from some common element naturalists however use such language only metaphorical sense they are far from meaning that during long course descent primordial organs any kind vertebræ the one case and legs the other have actually been modified into skulls jaws yet strong the appearance modification this nature having occurred that naturalists can hardly avoid employing language having this plain signification view these terms may used literally and the wonderful fact the jaws for instance crab retaining numerous characters which they would probably have retained through inheritance they had really been metamorphosed during long course descent from true legs from some simple appendage explained embryology has already been casually remarked that certain organs the individual which when mature become widely different and serve for different purposes are the embryo exactly alike the embryos also distinct animals within the same class are often strikingly similar better proof this cannot given than circumstance mentioned agassiz namely that having forgotten ticket the embryo some vertebrate animal cannot now tell whether that mammal bird reptile the vermiform larvæ moths flies beetles etc resemble each other much more closely than the mature insects but the case larvæ the embryos are active and have been adapted for special lines life trace the law embryonic resemblance sometimes lasts till rather late age thus birds the same genus and closely allied genera often resemble each other their first and second plumage see the spotted feathers the thrush group the cat tribe most the species are striped spotted lines and stripes can plainly distinguished the whelp the lion occasionally though rarely see something this kind plants thus the embryonic leaves the ulex furze and the first leaves the phyllodineous acaceas are pinnate divided like the ordinary leaves the leguminosæ the points structure which the embryos widely different animals the same class resemble each other often have direct relation their conditions existence cannot for instance suppose that the embryos the vertebrata the peculiar loop like course the arteries near the branchial slits are related similar conditions the young mammal which nourished the womb its mother the egg the bird which hatched nest and the spawn frog under water have more reason believe such relation than have believe that the same bones the hand man wing bat and fin porpoise are related similar conditions life one will suppose that the stripes the whelp lion the spots the young blackbird are any use these animals are related the conditions which they are exposed the case however different when animal during any part its embryonic career active and has provide for itself the period activity may come earlier later life but whenever comes the adaptation the larva its conditions life just perfect and beautiful the adult animal from such special adaptations the similarity the larvæ active embryos allied animals sometimes much obscured and cases could given the larvæ two species two groups species differing quite much even more from each other than their adult parents most cases however the larvæ though active still obey more less closely the law common embryonic resemblance cirripedes afford good instance this even the illustrious cuvier did not perceive that barnacle was certainly crustacean but glance the larva shows this the case unmistakeable manner again the two main divisions cirripedes the pedunculated and sessile which differ widely external appearance have larvæ all their several stages barely distinguishable the embryo the course development generally rises organisation use this expression though aware that hardly possible define clearly what meant the organisation being higher lower but one probably will dispute that the butterfly higher than the caterpillar some cases however the mature animal generally considered lower the scale than the larva with certain parasitic crustaceans refer once again cirripedes the larvæ the first stage have three pairs legs very simple single eye and probosciformed mouth with which they feed largely for they increase much size the second stage answering the chrysalis stage butterflies they have six pairs beautifully constructed natatory legs pair magnificent compound eyes and extremely complex antennæ but they have closed and imperfect mouth and cannot feed their function this stage search their well developed organs sense and reach their active powers swimming proper place which become attached and undergo their final metamorphosis when this completed they are fixed for life their legs are now converted into prehensile organs they again obtain well constructed mouth but they have antennæ and their two eyes are now reconverted into minute single and very simple eye spot this last and complete state cirripedes may considered either more highly more lowly organised than they were the larval condition but some genera the larvæ become developed either into hermaphrodites having the ordinary structure into what have called complemental males and the latter the development has assuredly been retrograde for the male mere sack which lives for short time and destitute mouth stomach other organ importance excepting for reproduction are much accustomed see differences structure between the embryo and the adult and likewise close similarity the embryos widely different animals within the same class that might led look these facts necessarily contingent some manner growth but there obvious reason why for instance the wing bat the fin porpoise should not have been sketched out with all the parts proper proportion soon any structure became visible the embryo and some whole groups animals and certain members other groups the embryo does not any period differ widely from the adult thus owen has remarked regard cuttle fish there metamorphosis the cephalopodic character manifested long before the parts the embryo are completed and again spiders there nothing worthy called metamorphosis the larvæ insects whether adapted the most diverse and active habits quite inactive being fed their parents placed the midst proper nutriment yet nearly all pass through similar worm like stage development but some few cases that aphis look the admirable drawings professor huxley the development this insect see trace the vermiform stage how then can explain these several facts embryology namely the very general but not universal difference structure between the embryo and the adult parts the same individual embryo which ultimately become very unlike and serve for diverse purposes being this early period growth alike embryos different species within the same class generally but not universally resembling each other the structure the embryo not being closely related its conditions existence except when the embryo becomes any period life active and has provide for itself the embryo apparently having sometimes higher organisation than the mature animal into which developed believe that all these facts can explained follows the view descent with modification commonly assumed perhaps from monstrosities often affecting the embryo very early period that slight variations necessarily appear equally early period but have little evidence this head indeed the evidence rather points the other way for notorious that breeders cattle horses and various fancy animals cannot positively tell until some time after the animal has been born what its merits form will ultimately turn out see this plainly our own children cannot always tell whether the child will tall short what its precise features will the question not what period life any variation has been caused but what period fully displayed the cause may have acted and believe generally has acted even before the embryo formed and the variation may due the male and female sexual elements having been affected the conditions which either parent their ancestors have been exposed nevertheless effect thus caused very early period even before the formation the embryo may appear late life when hereditary disease which appears old age alone has been communicated the offspring from the reproductive element one parent again when the horns cross bred cattle have been affected the shape the horns either parent for the welfare very young animal long remains its mother womb the egg long nourished and protected its parent must quite unimportant whether most its characters are fully acquired little earlier later life would not signify for instance bird which obtained its food best having long beak whether not assumed beak this particular length long was fed its parents hence conclude that quite possible that each the many successive modifications which each species has acquired its present structure may have supervened not very early period life and some direct evidence from our domestic animals supports this view but other cases quite possible that each successive modification most them may have appeared extremely early period have stated the first chapter that there some evidence render probable that whatever age any variation first appears the parent tends reappear corresponding age the offspring certain variations can only appear corresponding ages for instance peculiarities the caterpillar cocoon imago states the silk moth again the horns almost full grown cattle but further than this variations which for all that can see might have appeared earlier later life tend appear corresponding age the offspring and parent far from meaning that this invariably the case and could give good many cases variations taking the word the largest sense which have supervened earlier age the child than the parent these two principles their truth admitted will believe explain all the above specified leading facts embryology but first let look few analogous cases domestic varieties some authors who have written dogs maintain that the greyhound and bulldog though appearing different are really varieties most closely allied and have probably descended from the same wild stock hence was curious see how far their puppies differed from each other was told breeders that they differed just much their parents and this judging the eye seemed almost the case but actually measuring the old dogs and their six days old puppies found that the puppies had not nearly acquired their full amount proportional difference again was told that the foals cart and race horses differed much the full grown animals and this surprised greatly think probable that the difference between these two breeds has been wholly caused selection under domestication but having had careful measurements made the dam and three days old colt race and heavy cart horse find that the colts have means acquired their full amount proportional difference the evidence appears conclusive that the several domestic breeds pigeon have descended from one wild species compared young pigeons various breeds within twelve hours after being hatched carefully measured the proportions but will not here give details the beak width mouth length nostril and eyelid size feet and length leg the wild stock pouters fantails runts barbs dragons carriers and tumblers now some these birds when mature differ extraordinarily length and form beak that they would cannot doubt ranked distinct genera had they been natural productions but when the nestling birds these several breeds were placed row though most them could distinguished from each other yet their proportional differences the above specified several points were incomparably less than the full grown birds some characteristic points difference for instance that the width mouth could hardly detected the young but there was one remarkable exception this rule for the young the short faced tumbler differed from the young the wild rock pigeon and the other breeds all its proportions almost exactly much the adult state the two principles above given seem explain these facts regard the later embryonic stages our domestic varieties fanciers select their horses dogs and pigeons for breeding when they are nearly grown they are indifferent whether the desired qualities and structures have been acquired earlier later life the full grown animal possesses them and the cases just given more especially that pigeons seem show that the characteristic differences which give value each breed and which have been accumulated man selection have not generally first appeared early period life and have been inherited the offspring corresponding not early period but the case the short faced tumbler which when twelve hours old had acquired its proper proportions proves that this not the universal rule for here the characteristic differences must either have appeared earlier period than usual not the differences must have been inherited not the corresponding but earlier age now let apply these facts and the above two principles which latter though not proved true can shown some degree probable species state nature let take genus birds descended theory from some one parent species and which the several new species have become modified through natural selection accordance with their diverse habits then from the many slight successive steps variation having supervened rather late age and having been inherited corresponding age the young the new species our supposed genus will manifestly tend resemble each other much more closely than the adults just have seen the case pigeons may extend this view whole families even classes the fore limbs for instance which served legs the parent species may become long course modification adapted one descendant act hands another paddles another wings and the above two principles namely each successive modification supervening rather late age and being inherited corresponding late age the fore limbs the embryos the several descendants the parent species will still resemble each other closely for they will not have been modified but each individual new species the embryonic fore limbs will differ greatly from the fore limbs the mature animal the limbs the latter having undergone much modification rather late period life and having thus been converted into hands paddles wings whatever influence long continued exercise use the one hand and disuse the other may have modifying organ such influence will mainly affect the mature animal which has come its full powers activity and has gain its own living and the effects thus produced will inherited corresponding mature age whereas the young will remain unmodified modified lesser degree the effects use and disuse certain cases the successive steps variation might supervene from causes which are wholly ignorant very early period life each step might inherited earlier period than that which first appeared either case with the short faced tumbler the young embryo would closely resemble the mature parent form have seen that this the rule development certain whole groups animals with cuttle fish and spiders and with few members the great class insects with aphis with respect the final cause the young these cases not undergoing any metamorphosis closely resembling their parents from their earliest age can see that this would result from the two following contingencies firstly from the young during course modification carried for many generations having provide for their own wants very early stage development and secondly from their following exactly the same habits life with their parents for this case would indispensable for the existence the species that the child should modified very early age the same manner with its parents accordance with their similar habits some further explanation however the embryo not undergoing any metamorphosis perhaps requisite the other hand profited the young follow habits life any degree different from those their parent and consequently constructed slightly different manner then the principle inheritance corresponding ages the active young larvæ might easily rendered natural selection different any conceivable extent from their parents such differences might also become correlated with successive stages development that the larvæ the first stage might differ greatly from the larvæ the second stage have seen the case with cirripedes the adult might become fitted for sites habits which organs locomotion the senses etc would useless and this case the final metamorphosis would said retrograde all the organic beings extinct and recent which have ever lived this earth have classed together and all have been connected the finest gradations the best indeed our collections were nearly perfect the only possible arrangement would genealogical descent being view the hidden bond connexion which naturalists have been seeking under the term the natural system this view can understand how that the eyes most naturalists the structure the embryo even more important for classification than that the adult for the embryo the animal its less modified state and far reveals the structure its progenitor two groups animal however much they may present differ from each other structure and habits they pass through the same similar embryonic stages may feel assured that they have both descended from the same nearly similar parents and are therefore that degree closely related thus community embryonic structure reveals community descent will reveal this community descent however much the structure the adult may have been modified and obscured have seen for instance that cirripedes can once recognised their larvæ belonging the great class crustaceans the embryonic state each species and group species partially shows the structure their less modified ancient progenitors can clearly see why ancient and extinct forms life should resemble the embryos their descendants our existing species agassiz believes this law nature but bound confess that only hope see the law hereafter proved true can proved true those cases alone which the ancient state now supposed represented many embryos has not been obliterated either the successive variations long course modification having supervened very early age the variations having been inherited earlier period than that which they first appeared should also borne mind that the supposed law resemblance ancient forms life the embryonic stages recent forms may true but yet owing the geological record not extending far enough back time may remain for long period for ever incapable demonstration thus seems the leading facts embryology which are second importance none natural history are explained the principle slight modifications not appearing the many descendants from some one ancient progenitor very early period the life each though perhaps caused the earliest and being inherited corresponding not early period embryology rises greatly interest when thus look the embryo picture more less obscured the common parent form each great class animals rudimentary atrophied aborted organs organs parts this strange condition bearing the stamp inutility are extremely common throughout nature for instance rudimentary mammæ are very general the males mammals presume that the bastard wing birds may safely considered digit rudimentary state very many snakes one lobe the lungs rudimentary other snakes there are rudiments the pelvis and hind limbs some the cases rudimentary organs are extremely curious for instance the presence teeth foetal whales which when grown have not tooth their heads and the presence teeth which never cut through the gums the upper jaws our unborn calves has even been stated good authority that rudiments teeth can detected the beaks certain embryonic birds nothing can plainer than that wings are formed for flight yet how many insects see wings reduced size utterly incapable flight and not rarely lying under wing cases firmly soldered together the meaning rudimentary organs often quite unmistakeable for instance there are beetles the same genus and even the same species resembling each other most closely all respects one which will have full sized wings and another mere rudiments membrane and here impossible doubt that the rudiments represent wings rudimentary organs sometimes retain their potentiality and are merely not developed this seems the case with the mammæ male mammals for many instances are record these organs having become well developed full grown males and having secreted milk again there are normally four developed and two rudimentary teats the udders the genus bos but our domestic cows the two sometimes become developed and give milk individual plants the same species the petals sometimes occur mere rudiments and sometimes well developed state plants with separated sexes the male flowers often have rudiment pistil and kölreuter found that crossing such male plants with hermaphrodite species the rudiment the pistil the hybrid offspring was much increased size and this shows that the rudiment and the perfect pistil are essentially alike nature organ serving for two purposes may become rudimentary utterly aborted for one even the more important purpose and remain perfectly efficient for the other thus plants the office the pistil allow the pollen tubes reach the ovules protected the ovarium its base the pistil consists stigma supported the style but some compositæ the male florets which course cannot fecundated have pistil which rudimentary state for not crowned with stigma but the style remains well developed and clothed with hairs other compositæ for the purpose brushing the pollen out the surrounding anthers again organ may become rudimentary for its proper purpose and used for distinct object certain fish the swim bladder seems rudimentary for its proper function giving buoyancy but has become converted into nascent breathing organ lung other similar instances could given rudimentary organs the individuals the same species are very liable vary degree development and other respects moreover closely allied species the degree which the same organ has been rendered rudimentary occasionally differs much this latter fact well exemplified the state the wings the female moths certain groups rudimentary organs may utterly aborted and this implies that find animal plant trace organ which analogy would lead expect find and which occasionally found monstrous individuals the species thus the snapdragon antirrhinum generally not find rudiment fifth stamen but this may sometimes seen tracing the homologies the same part different members class nothing more common more necessary than the use and discovery rudiments this well shown the drawings given owen the bones the leg the horse and rhinoceros important fact that rudimentary organs such teeth the upper jaws whales and ruminants can often detected the embryo but afterwards wholly disappear also believe universal rule that rudimentary part organ greater size relatively the adjoining parts the embryo than the adult that the organ this early age less rudimentary even cannot said any degree rudimentary hence also rudimentary organ the adult often said have retained its embryonic condition have now given the leading facts with respect rudimentary organs reflecting them every one must struck with astonishment for the same reasoning power which tells plainly that most parts and organs are exquisitely adapted for certain purposes tells with equal plainness that these rudimentary atrophied organs are imperfect and useless works natural history rudimentary organs are generally said have been created for the sake symmetry order complete the scheme nature but this seems explanation merely restatement the fact would thought sufficient say that because planets revolve elliptic courses round the sun satellites follow the same course round the planets for the sake symmetry and complete the scheme nature eminent physiologist accounts for the presence rudimentary organs supposing that they serve excrete matter excess injurious the system but can suppose that the minute papilla which often represents the pistil male flowers and which formed merely cellular tissue can thus act can suppose that the formation rudimentary teeth which are subsequently absorbed can any service the rapidly growing embryonic calf the excretion precious phosphate lime when man fingers have been amputated imperfect nails sometimes appear the stumps could soon believe that these vestiges nails have appeared not from unknown laws growth but order excrete horny matter that the rudimentary nails the fin the manatee were formed for this purpose view descent with modification the origin rudimentary organs simple have plenty cases rudimentary organs our domestic productions the stump tail tailless breeds the vestige ear earless breeds the reappearance minute dangling horns hornless breeds cattle more especially according youatt young animals and the state the whole flower the cauliflower often see rudiments various parts monsters but doubt whether any these cases throw light the origin rudimentary organs state nature further than showing that rudiments can produced for doubt whether species under nature ever undergo abrupt changes believe that disuse has been the main agency that has led successive generations the gradual reduction various organs until they have become rudimentary the case the eyes animals inhabiting dark caverns and the wings birds inhabiting oceanic islands which have seldom been forced take flight and have ultimately lost the power flying again organ useful under certain conditions might become injurious under others with the wings beetles living small and exposed islands and this case natural selection would continue slowly reduce the organ until was rendered harmless and rudimentary any change function which can effected insensibly small steps within the power natural selection that organ rendered during changed habits life useless injurious for one purpose might easily modified and used for another purpose organ might retained for one alone its former functions organ when rendered useless may well variable for its variations cannot checked natural selection whatever period life disuse selection reduces organ and this will generally when the being has come maturity and its full powers action the principle inheritance corresponding ages will reproduce the organ its reduced state the same age and consequently will seldom affect reduce the embryo thus can understand the greater relative size rudimentary organs the embryo and their lesser relative size the adult but each step the process reduction were inherited not the corresponding age but extremely early period life have good reason believe possible the rudimentary part would tend wholly lost and should have case complete abortion the principle also economy explained former chapter which the materials forming any part structure not useful the possessor will saved far possible will probably often come into play and this will tend cause the entire obliteration rudimentary organ the presence rudimentary organs thus due the tendency every part the organisation which has long existed inherited can understand the genealogical view classification how that systematists have found rudimentary parts useful even sometimes more useful than parts high physiological importance rudimentary organs may compared with the letters word still retained the spelling but become useless the pronunciation but which serve clue seeking for its derivation the view descent with modification may conclude that the existence organs rudimentary imperfect and useless condition quite aborted far from presenting strange difficulty they assuredly the ordinary doctrine creation might even have been anticipated and can accounted for the laws inheritance summary this chapter have attempted show that the subordination group group all organisms throughout all time that the nature the relationship which all living and extinct beings are united complex radiating and circuitous lines affinities into one grand system the rules followed and the difficulties encountered naturalists their classifications the value set upon characters constant and prevalent whether high vital importance the most trifling importance rudimentary organs importance the wide opposition value between analogical adaptive characters and characters true affinity and other such rules all naturally follow the view the common parentage those forms which are considered naturalists allied together with their modification through natural selection with its contingencies extinction and divergence character considering this view classification should borne mind that the element descent has been universally used ranking together the sexes ages and acknowledged varieties the same species however different they may structure extend the use this element descent the only certainly known cause similarity organic beings shall understand what meant the natural system genealogical its attempted arrangement with the grades acquired difference marked the terms varieties species genera families orders and classes this same view descent with modification all the great facts morphology become intelligible whether look the same pattern displayed the homologous organs whatever purpose applied the different species class the homologous parts constructed the same pattern each individual animal and plant the principle successive slight variations not necessarily generally supervening very early period life and being inherited corresponding period can understand the great leading facts embryology namely the resemblance individual embryo the homologous parts which when matured will become widely different from each other structure and function and the resemblance different species class the homologous parts organs though fitted the adult members for purposes different possible larvæ are active embryos which have become specially modified relation their habits life through the principle modifications being inherited corresponding ages this same principle and bearing mind that when organs are reduced size either from disuse selection will generally that period life when the being has provide for its own wants and bearing mind how strong the principle inheritance the occurrence rudimentary organs and their final abortion present inexplicable difficulties the contrary their presence might have been even anticipated the importance embryological characters and rudimentary organs classification intelligible the view that arrangement only far natural genealogical finally the several classes facts which have been considered this chapter seem proclaim plainly that the innumerable species genera and families organic beings with which this world peopled have all descended each within its own class group from common parents and have all been modified the course descent that should without hesitation adopt this view even were unsupported other facts arguments chapter xiv recapitulation and conclusion recapitulation the difficulties the theory natural selection recapitulation the general and special circumstances its favour causes the general belief the immutability species how far the theory natural selection may extended effects its adoption the study natural history concluding remarks this whole volume one long argument may convenient the reader have the leading facts and inferences briefly recapitulated that many and grave objections may advanced against the theory descent with modification through natural selection not deny have endeavoured give them their full force nothing first can appear more difficult believe than that the more complex organs and instincts should have been perfected not means superior though analogous with human reason but the accumulation innumerable slight variations each good for the individual possessor nevertheless this difficulty though appearing our imagination insuperably great cannot considered real admit the following propositions namely that gradations the perfection any organ instinct which may consider either now exist could have existed each good its kind that all organs and instincts are ever slight degree variable and lastly that there struggle for existence leading the preservation each profitable deviation structure instinct the truth these propositions cannot think disputed doubt extremely difficult even conjecture what gradations many structures have been perfected more especially amongst broken and failing groups organic beings but see many strange gradations nature proclaimed the canon natura non facit saltum that ought extremely cautious saying that any organ instinct any whole being could not have arrived its present state many graduated steps there are must admitted cases special difficulty the theory natural selection and one the most curious these the existence two three defined castes workers sterile females the same community ants but have attempted show how this difficulty can mastered with respect the almost universal sterility species when first crossed which forms remarkable contrast with the almost universal fertility varieties when crossed must refer the reader the recapitulation the facts given the end the eighth chapter which seem conclusively show that this sterility more special endowment than the incapacity two trees grafted together but that incidental constitutional differences the reproductive systems the intercrossed species see the truth this conclusion the vast difference the result when the same two species are crossed reciprocally that when one species first used the father and then the mother the fertility varieties when intercrossed and their mongrel offspring cannot considered universal nor their very general fertility surprising when remember that not likely that either their constitutions their reproductive systems should have been profoundly modified moreover most the varieties which have been experimentised have been produced under domestication and domestication apparently tends eliminate sterility ought not expect also produce sterility the sterility hybrids very different case from that first crosses for their reproductive organs are more less functionally impotent whereas first crosses the organs both sides are perfect condition continually see that organisms all kinds are rendered some degree sterile from their constitutions having been disturbed slightly different and new conditions life need not feel surprise hybrids being some degree sterile for their constitutions can hardly fail have been disturbed from being compounded two distinct organisations this parallelism supported another parallel but directly opposite class facts namely that the vigour and fertility all organic beings are increased slight changes their conditions life and that the offspring slightly modified forms varieties acquire from being crossed increased vigour and fertility that the one hand considerable changes the conditions life and crosses between greatly modified forms lessen fertility and the other hand lesser changes the conditions life and crosses between less modified forms increase fertility turning geographical distribution the difficulties encountered the theory descent with modification are grave enough all the individuals the same species and all the species the same genus even higher group must have descended from common parents and therefore however distant and isolated parts the world they are now found they must the course successive generations have passed from some one part the others are often wholly unable even conjecture how this could have been effected yet have reason believe that some species have retained the same specific form for very long periods enormously long measured years too much stress ought not laid the occasional wide diffusion the same species for during very long periods time there will always good chance for wide migration many means broken interrupted range may often accounted for the extinction the species the intermediate regions cannot denied that are yet very ignorant the full extent the various climatal and geographical changes which have affected the earth during modern periods and such changes will obviously have greatly facilitated migration example have attempted show how potent has been the influence the glacial period the distribution both the same and representative species throughout the world are yet profoundly ignorant the many occasional means transport with respect distinct species the same genus inhabiting very distant and isolated regions the process modification has necessarily been slow all the means migration will have been possible during very long period and consequently the difficulty the wide diffusion species the same genus some degree lessened the theory natural selection interminable number intermediate forms must have existed linking together all the species each group gradations fine our present varieties may asked why not see these linking forms all around why are not all organic beings blended together inextricable chaos with respect existing forms should remember that have right expect excepting rare cases discover directly connecting links between them but only between each and some extinct and supplanted form even wide area which has during long period remained continuous and which the climate and other conditions life change insensibly going from district occupied one species into another district occupied closely allied species have just right expect often find intermediate varieties the intermediate zone for have reason believe that only few species are undergoing change any one period and all changes are slowly effected have also shown that the intermediate varieties which will first probably exist the intermediate zones will liable supplanted the allied forms either hand and the latter from existing greater numbers will generally modified and improved quicker rate than the intermediate varieties which exist lesser numbers that the intermediate varieties will the long run supplanted and exterminated this doctrine the extermination infinitude connecting links between the living and extinct inhabitants the world and each successive period between the extinct and still older species why not every geological formation charged with such links why does not every collection fossil remains afford plain evidence the gradation and mutation the forms life meet with such evidence and this the most obvious and forcible the many objections which may urged against theory why again whole groups allied species appear though certainly they often falsely appear have come suddenly the several geological stages why not find great piles strata beneath the silurian system stored with the remains the progenitors the silurian groups fossils for certainly theory such strata must somewhere have been deposited these ancient and utterly unknown epochs the world history can answer these questions and grave objections only the supposition that the geological record far more imperfect than most geologists believe cannot objected that there has not been time sufficient for any amount organic change for the lapse time has been great utterly inappreciable the human intellect the number specimens all our museums absolutely nothing compared with the countless generations countless species which certainly have existed should not able recognise species the parent any one more species were examine them ever closely unless likewise possessed many the intermediate links between their past parent and present states and these many links could hardly ever expect discover owing the imperfection the geological record numerous existing doubtful forms could named which are probably varieties but who will pretend that future ages many fossil links will discovered that naturalists will able decide the common view whether not these doubtful forms are varieties long most the links between any two species are unknown any one link intermediate variety discovered will simply classed another and distinct species only small portion the world has been geologically explored only organic beings certain classes can preserved fossil condition least any great number widely ranging species vary most and varieties are often first local both causes rendering the discovery intermediate links less likely local varieties will not spread into other and distant regions until they are considerably modified and improved and when they spread discovered geological formation they will appear suddenly created there and will simply classed new species most formations have been intermittent their accumulation and their duration inclined believe has been shorter than the average duration specific forms successive formations are separated from each other enormous blank intervals time for fossiliferous formations thick enough resist future degradation can accumulated only where much sediment deposited the subsiding bed the sea during the alternate periods elevation and stationary level the record will blank during these latter periods there will probably more variability the forms life during periods subsidence more extinction with respect the absence fossiliferous formations beneath the lowest silurian strata can only recur the hypothesis given the ninth chapter that the geological record imperfect all will admit but that imperfect the degree which require few will inclined admit look long enough intervals time geology plainly declares that all species have changed and they have changed the manner which theory requires for they have changed slowly and graduated manner clearly see this the fossil remains from consecutive formations invariably being much more closely related each other than are the fossils from formations distant from each other time such the sum the several chief objections and difficulties which may justly urged against theory and have now briefly recapitulated the answers and explanations which can given them have felt these difficulties far too heavily during many years doubt their weight but deserves especial notice that the more important objections relate questions which are confessedly ignorant nor know how ignorant are not know all the possible transitional gradations between the simplest and the most perfect organs cannot pretended that know all the varied means distribution during the long lapse years that know how imperfect the geological record grave these several difficulties are judgment they not overthrow the theory descent with modification now let turn the other side the argument under domestication see much variability this seems mainly due the reproductive system being eminently susceptible changes the conditions life that this system when not rendered impotent fails reproduce offspring exactly like the parent form variability governed many complex laws correlation growth use and disuse and the direct action the physical conditions life there much difficulty ascertaining how much modification our domestic productions have undergone but may safely infer that the amount has been large and that modifications can inherited for long periods long the conditions life remain the same have reason believe that modification which has already been inherited for many generations may continue inherited for almost infinite number generations the other hand have evidence that variability when has once come into play does not wholly cease for new varieties are still occasionally produced our most anciently domesticated productions man does not actually produce variability only unintentionally exposes organic beings new conditions life and then nature acts the organisation and causes variability but man can and does select the variations given him nature and thus accumulate them any desired manner thus adapts animals and plants for his own benefit pleasure may this methodically may unconsciously preserving the individuals most useful him the time without any thought altering the breed certain that can largely influence the character breed selecting each successive generation individual differences slight quite inappreciable uneducated eye this process selection has been the great agency the production the most distinct and useful domestic breeds that many the breeds produced man have large extent the character natural species shown the inextricable doubts whether very many them are varieties aboriginal species there obvious reason why the principles which have acted efficiently under domestication should not have acted under nature the preservation favoured individuals and races during the constantly recurrent struggle for existence see the most powerful and ever acting means selection the struggle for existence inevitably follows from the high geometrical ratio increase which common all organic beings this high rate increase proved calculation the effects succession peculiar seasons and the results naturalisation explained the third chapter more individuals are born than can possibly survive grain the balance will determine which individual shall live and which shall die which variety species shall increase number and which shall decrease finally become extinct the individuals the same species come all respects into the closest competition with each other the struggle will generally most severe between them will almost equally severe between the varieties the same species and next severity between the species the same genus but the struggle will often very severe between beings most remote the scale nature the slightest advantage one being any age during any season over those with which comes into competition better adaptation however slight degree the surrounding physical conditions will turn the balance with animals having separated sexes there will most cases struggle between the males for possession the females the most vigorous individuals those which have most successfully struggled with their conditions life will generally leave most progeny but success will often depend having special weapons means defence the charms the males and the slightest advantage will lead victory geology plainly proclaims that each land has undergone great physical changes might have expected that organic beings would have varied under nature the same way they generally have varied under the changed conditions domestication and there any variability under nature would unaccountable fact natural selection had not come into play has often been asserted but the assertion quite incapable proof that the amount variation under nature strictly limited quantity man though acting external characters alone and often capriciously can produce within short period great result adding mere individual differences his domestic productions and every one admits that there are least individual differences species under nature but besides such differences all naturalists have admitted the existence varieties which they think sufficiently distinct worthy record systematic works one can draw any clear distinction between individual differences and slight varieties between more plainly marked varieties and sub species and species let observed how naturalists differ the rank which they assign the many representative forms europe and north america then have under nature variability and powerful agent always ready act and select why should doubt that variations any way useful beings under their excessively complex relations life would preserved accumulated and inherited why man can patience select variations most useful himself should nature fail selecting variations useful under changing conditions life her living products what limit can put this power acting during long ages and rigidly scrutinising the whole constitution structure and habits each creature favouring the good and rejecting the bad can see limit this power slowly and beautifully adapting each form the most complex relations life the theory natural selection even looked further than this seems itself probable have already recapitulated fairly could the opposed difficulties and objections now let turn the special facts and arguments favour the theory the view that species are only strongly marked and permanent varieties and that each species first existed variety can see why that line demarcation can drawn between species commonly supposed have been produced special acts creation and varieties which are acknowledged have been produced secondary laws this same view can understand how that each region where many species genus have been produced and where they now flourish these same species should present many varieties for where the manufactory species has been active might expect general rule find still action and this the case varieties incipient species moreover the species the larger genera which afford the greater number varieties incipient species retain certain degree the character varieties for they differ from each other less amount difference than the species smaller genera the closely allied species also the larger genera apparently have restricted ranges and they are clustered little groups round other species which respects they resemble varieties these are strange relations the view each species having been independently created but are intelligible all species first existed varieties each species tends its geometrical ratio reproduction increase inordinately number and the modified descendants each species will enabled increase much the more they become more diversified habits and structure enabled seize many and widely different places the economy nature there will constant tendency natural selection preserve the most divergent offspring any one species hence during long continued course modification the slight differences characteristic varieties the same species tend augmented into the greater differences characteristic species the same genus new and improved varieties will inevitably supplant and exterminate the older less improved and intermediate varieties and thus species are rendered large extent defined and distinct objects dominant species belonging the larger groups tend give birth new and dominant forms that each large group tends become still larger and the same time more divergent character but all groups cannot thus succeed increasing size for the world would not hold them the more dominant groups beat the less dominant this tendency the large groups increasing size and diverging character together with the almost inevitable contingency much extinction explains the arrangement all the forms life groups subordinate groups all within few great classes which now see everywhere around and which has prevailed throughout all time this grand fact the grouping all organic beings seems utterly inexplicable the theory creation natural selection acts solely accumulating slight successive favourable variations can produce great sudden modification can act only very short and slow steps hence the canon natura non facit saltum which every fresh addition our knowledge tends make more strictly correct this theory simply intelligible can plainly see why nature prodigal variety though niggard innovation but why this should law nature each species has been independently created man can explain many other facts are seems explicable this theory how strange that bird under the form woodpecker should have been created prey insects the ground that upland geese which never rarely swim should have been created with webbed feet that thrush should have been created dive and feed sub aquatic insects and that petrel should have been created with habits and structure fitting for the life auk grebe and endless other cases but the view each species constantly trying increase number with natural selection always ready adapt the slowly varying descendants each any unoccupied ill occupied place nature these facts cease strange perhaps might even have been anticipated natural selection acts competition adapts the inhabitants each country only relation the degree perfection their associates that need feel surprise the inhabitants any one country although the ordinary view supposed have been specially created and adapted for that country being beaten and supplanted the naturalised productions from another land nor ought marvel all the contrivances nature not far can judge absolutely perfect and some them abhorrent our ideas fitness need not marvel the sting the bee causing the bee own death drones being produced such vast numbers for one single act and being then slaughtered their sterile sisters the astonishing waste pollen our fir trees the instinctive hatred the queen bee for her own fertile daughters ichneumonidæ feeding within the live bodies caterpillars and other such cases the wonder indeed the theory natural selection that more cases the want absolute perfection have not been observed the complex and little known laws governing variation are the same far can see with the laws which have governed the production called specific forms both cases physical conditions seem have produced but little direct effect yet when varieties enter any zone they occasionally assume some the characters the species proper that zone both varieties and species use and disuse seem have produced some effect for difficult resist this conclusion when look for instance the logger headed duck which has wings incapable flight nearly the same condition the domestic duck when look the burrowing tucutucu which occasionally blind and then certain moles which are habitually blind and have their eyes covered with skin when look the blind animals inhabiting the dark caves america and europe both varieties and species correlation growth seems have played most important part that when one part has been modified other parts are necessarily modified both varieties and species reversions long lost characters occur how inexplicable the theory creation the occasional appearance stripes the shoulder and legs the several species the horse genus and their hybrids how simply this fact explained believe that these species have descended from striped progenitor the same manner the several domestic breeds pigeon have descended from the blue and barred rock pigeon the ordinary view each species having been independently created why should the specific characters those which the species the same genus differ from each other more variable than the generic characters which they all agree why for instance should the colour flower more likely vary any one species genus the other species supposed have been created independently have differently coloured flowers than all the species the genus have the same coloured flowers species are only well marked varieties which the characters have become high degree permanent can understand this fact for they have already varied since they branched off from common progenitor certain characters which they have come specifically distinct from each other and therefore these same characters would more likely still variable than the generic characters which have been inherited without change for enormous period inexplicable the theory creation why part developed very unusual manner any one species genus and therefore may naturally infer great importance the species should eminently liable variation but view this part has undergone since the several species branched off from common progenitor unusual amount variability and modification and therefore might expect this part generally still variable but part may developed the most unusual manner like the wing bat and yet not more variable than any other structure the part common many subordinate forms that has been inherited for very long period for this case will have been rendered constant long continued natural selection glancing instincts marvellous some are they offer greater difficulty than does corporeal structure the theory the natural selection successive slight but profitable modifications can thus understand why nature moves graduated steps endowing different animals the same class with their several instincts have attempted show how much light the principle gradation throws the admirable architectural powers the hive bee habit doubt sometimes comes into play modifying instincts but certainly not indispensable see the case neuter insects which leave progeny inherit the effects long continued habit the view all the species the same genus having descended from common parent and having inherited much common can understand how that allied species when placed under considerably different conditions life yet should follow nearly the same instincts why the thrush south america for instance lines her nest with mud like our british species the view instincts having been slowly acquired through natural selection need not marvel some instincts being apparently not perfect and liable mistakes and many instincts causing other animals suffer species only well marked and permanent varieties can once see why their crossed offspring should follow the same complex laws their degrees and kinds resemblance their parents being absorbed into each other successive crosses and other such points the crossed offspring acknowledged varieties the other hand these would strange facts species have been independently created and varieties have been produced secondary laws admit that the geological record imperfect extreme degree then such facts the record gives support the theory descent with modification new species have come the stage slowly and successive intervals and the amount change after equal intervals time widely different different groups the extinction species and whole groups species which has played conspicuous part the history the organic world almost inevitably follows the principle natural selection for old forms will supplanted new and improved forms neither single species nor groups species reappear when the chain ordinary generation has once been broken the gradual diffusion dominant forms with the slow modification their descendants causes the forms life after long intervals time appear they had changed simultaneously throughout the world the fact the fossil remains each formation being some degree intermediate character between the fossils the formations above and below simply explained their intermediate position the chain descent the grand fact that all extinct organic beings belong the same system with recent beings falling either into the same into intermediate groups follows from the living and the extinct being the offspring common parents the groups which have descended from ancient progenitor have generally diverged character the progenitor with its early descendants will often intermediate character comparison with its later descendants and thus can see why the more ancient fossil the oftener stands some degree intermediate between existing and allied groups recent forms are generally looked being some vague sense higher than ancient and extinct forms and they are far higher the later and more improved forms have conquered the older and less improved organic beings the struggle for life lastly the law the long endurance allied forms the same continent marsupials australia edentata america and other such cases intelligible for within confined country the recent and the extinct will naturally allied descent looking geographical distribution admit that there has been during the long course ages much migration from one part the world another owing former climatal and geographical changes and the many occasional and unknown means dispersal then can understand the theory descent with modification most the great leading facts distribution can see why there should striking parallelism the distribution organic beings throughout space and their geological succession throughout time for both cases the beings have been connected the bond ordinary generation and the means modification have been the same see the full meaning the wonderful fact which must have struck every traveller namely that the same continent under the most diverse conditions under heat and cold mountain and lowland deserts and marshes most the inhabitants within each great class are plainly related for they will generally descendants the same progenitors and early colonists this same principle former migration combined most cases with modification can understand the aid the glacial period the identity some few plants and the close alliance many others the most distant mountains under the most different climates and likewise the close alliance some the inhabitants the sea the northern and southern temperate zones though separated the whole intertropical ocean although two areas may present the same physical conditions life need feel surprise their inhabitants being widely different they have been for long period completely separated from each other for the relation organism organism the most important all relations and the two areas will have received colonists from some third source from each other various periods and different proportions the course modification the two areas will inevitably different this view migration with subsequent modification can see why oceanic islands should inhabited few species but these that many should peculiar can clearly see why those animals which cannot cross wide spaces ocean frogs and terrestrial mammals should not inhabit oceanic islands and why the other hand new and peculiar species bats which can traverse the ocean should often found islands far distant from any continent such facts the presence peculiar species bats and the absence all other mammals oceanic islands are utterly inexplicable the theory independent acts creation the existence closely allied representative species any two areas implies the theory descent with modification that the same parents formerly inhabited both areas and almost invariably find that wherever many closely allied species inhabit two areas some identical species common both still exist wherever many closely allied yet distinct species occur many doubtful forms and varieties the same species likewise occur rule high generality that the inhabitants each area are related the inhabitants the nearest source whence immigrants might have been derived see this nearly all the plants and animals the galapagos archipelago juan fernandez and the other american islands being related the most striking manner the plants and animals the neighbouring american mainland and those the cape verde archipelago and other african islands the african mainland must admitted that these facts receive explanation the theory creation the fact have seen that all past and present organic beings constitute one grand natural system with group subordinate group and with extinct groups often falling between recent groups intelligible the theory natural selection with its contingencies extinction and divergence character these same principles see how that the mutual affinities the species and genera within each class are complex and circuitous see why certain characters are far more serviceable than others for classification why adaptive characters though paramount importance the being are hardly any importance classification why characters derived from rudimentary parts though service the being are often high classificatory value and why embryological characters are the most valuable all the real affinities all organic beings are due inheritance community descent the natural system genealogical arrangement which have discover the lines descent the most permanent characters however slight their vital importance may the framework bones being the same the hand man wing bat fin the porpoise and leg the horse the same number vertebræ forming the neck the giraffe and the elephant and innumerable other such facts once explain themselves the theory descent with slow and slight successive modifications the similarity pattern the wing and leg bat though used for such different purpose the jaws and legs crab the petals stamens and pistils flower likewise intelligible the view the gradual modification parts organs which were alike the early progenitor each class the principle successive variations not always supervening early age and being inherited corresponding not early period life can clearly see why the embryos mammals birds reptiles and fishes should closely alike and should unlike the adult forms may cease marvelling the embryo air breathing mammal bird having branchial slits and arteries running loops like those fish which has breathe the air dissolved water the aid well developed branchiæ disuse aided sometimes natural selection will often tend reduce organ when has become useless changed habits under changed conditions life and can clearly understand this view the meaning rudimentary organs but disuse and selection will generally act each creature when has come maturity and has play its full part the struggle for existence and will thus have little power acting organ during early life hence the organ will not much reduced rendered rudimentary this early age the calf for instance has inherited teeth which never cut through the gums the upper jaw from early progenitor having well developed teeth and may believe that the teeth the mature animal were reduced during successive generations disuse the tongue and palate having been fitted natural selection browse without their aid whereas the calf the teeth have been left untouched selection disuse and the principle inheritance corresponding ages have been inherited from remote period the present day the view each organic being and each separate organ having been specially created how utterly inexplicable that parts like the teeth the embryonic calf like the shrivelled wings under the soldered wing covers some beetles should thus frequently bear the plain stamp inutility nature may said have taken pains reveal rudimentary organs and homologous structures her scheme modification which seems that wilfully will not understand have now recapitulated the chief facts and considerations which have thoroughly convinced that species have changed and are still slowly changing the preservation and accumulation successive slight favourable variations why may asked have all the most eminent living naturalists and geologists rejected this view the mutability species cannot asserted that organic beings state nature are subject variation cannot proved that the amount variation the course long ages limited quantity clear distinction has been can drawn between species and well marked varieties cannot maintained that species when intercrossed are invariably sterile and varieties invariably fertile that sterility special endowment and sign creation the belief that species were immutable productions was almost unavoidable long the history the world was thought short duration and now that have acquired some idea the lapse time are too apt assume without proof that the geological record perfect that would have afforded plain evidence the mutation species they had undergone mutation but the chief cause our natural unwillingness admit that one species has given birth other and distinct species that are always slow admitting any great change which not see the intermediate steps the difficulty the same that felt many geologists when lyell first insisted that long lines inland cliffs had been formed and great valleys excavated the slow action the coast waves the mind cannot possibly grasp the full meaning the term hundred million years cannot add and perceive the full effects many slight variations accumulated during almost infinite number generations although fully convinced the truth the views given this volume under the form abstract means expect convince experienced naturalists whose minds are stocked with multitude facts all viewed during long course years from point view directly opposite mine easy hide our ignorance under such expressions the plan creation unity design etc and think that give explanation when only restate fact any one whose disposition leads him attach more weight unexplained difficulties than the explanation certain number facts will certainly reject theory few naturalists endowed with much flexibility mind and who have already begun doubt the immutability species may influenced this volume but look with confidence the future young and rising naturalists who will able view both sides the question with impartiality whoever led believe that species are mutable will good service conscientiously expressing his conviction for only thus can the load prejudice which this subject overwhelmed removed several eminent naturalists have late published their belief that multitude reputed species each genus are not real species but that other species are real that have been independently created this seems strange conclusion arrive they admit that multitude forms which till lately they themselves thought were special creations and which are still thus looked the majority naturalists and which consequently have every external characteristic feature true species they admit that these have been produced variation but they refuse extend the same view other and very slightly different forms nevertheless they not pretend that they can define even conjecture which are the created forms life and which are those produced secondary laws they admit variation vera causa one case they arbitrarily reject another without assigning any distinction the two cases the day will come when this will given curious illustration the blindness preconceived opinion these authors seem more startled miraculous act creation than ordinary birth but they really believe that innumerable periods the earth history certain elemental atoms have been commanded suddenly flash into living tissues they believe that each supposed act creation one individual many were produced were all the infinitely numerous kinds animals and plants created eggs seed full grown and the case mammals were they created bearing the false marks nourishment from the mother womb although naturalists very properly demand full explanation every difficulty from those who believe the mutability species their own side they ignore the whole subject the first appearance species what they consider reverent silence may asked how far extend the doctrine the modification species the question difficult answer because the more distinct the forms are which may consider much the arguments fall away force but some arguments the greatest weight extend very far all the members whole classes can connected together chains affinities and all can classified the same principle groups subordinate groups fossil remains sometimes tend fill very wide intervals between existing orders organs rudimentary condition plainly show that early progenitor had the organ fully developed state and this some instances necessarily implies enormous amount modification the descendants throughout whole classes various structures are formed the same pattern and embryonic age the species closely resemble each other therefore cannot doubt that the theory descent with modification embraces all the members the same class believe that animals have descended from most only four five progenitors and plants from equal lesser number analogy would lead one step further namely the belief that all animals and plants have descended from some one prototype but analogy may deceitful guide nevertheless all living things have much common their chemical composition their germinal vesicles their cellular structure and their laws growth and reproduction see this even trifling circumstance that the same poison often similarly affects plants and animals that the poison secreted the gall fly produces monstrous growths the wild rose oak tree therefore should infer from analogy that probably all the organic beings which have ever lived this earth have descended from some one primordial form into which life was first breathed when the views entertained this volume the origin species when analogous views are generally admitted can dimly foresee that there will considerable revolution natural history systematists will able pursue their labours present but they will not incessantly haunted the shadowy doubt whether this that form essence species this feel sure and speak after experience will slight relief the endless disputes whether not some fifty species british brambles are true species will cease systematists will have only decide not that this will easy whether any form sufficiently constant and distinct from other forms capable definition and definable whether the differences sufficiently important deserve specific name this latter point will become far more essential consideration than present for differences however slight between any two forms not blended intermediate gradations are looked most naturalists sufficient raise both forms the rank species hereafter shall compelled acknowledge that the only distinction between species and well marked varieties that the latter are known believed connected the present day intermediate gradations whereas species were formerly thus connected hence without quite rejecting the consideration the present existence intermediate gradations between any two forms shall led weigh more carefully and value higher the actual amount difference between them quite possible that forms now generally acknowledged merely varieties may hereafter thought worthy specific names with the primrose and cowslip and this case scientific and common language will come into accordance short shall have treat species the same manner those naturalists treat genera who admit that genera are merely artificial combinations made for convenience this may not cheering prospect but shall least freed from the vain search for the undiscovered and undiscoverable essence the term species the other and more general departments natural history will rise greatly interest the terms used naturalists affinity relationship community type paternity morphology adaptive characters rudimentary and aborted organs etc will cease metaphorical and will have plain signification when longer look organic being savage looks ship something wholly beyond his comprehension when regard every production nature one which has had history when contemplate every complex structure and instinct the summing many contrivances each useful the possessor nearly the same way when look any great mechanical invention the summing the labour the experience the reason and even the blunders numerous workmen when thus view each organic being how far more interesting speak from experience will the study natural history become grand and almost untrodden field inquiry will opened the causes and laws variation correlation growth the effects use and disuse the direct action external conditions and forth the study domestic productions will rise immensely value new variety raised man will far more important and interesting subject for study than one more species added the infinitude already recorded species our classifications will come far they can made genealogies and will then truly give what may called the plan creation the rules for classifying will doubt become simpler when have definite object view possess pedigrees armorial bearings and have discover and trace the many diverging lines descent our natural genealogies characters any kind which have long been inherited rudimentary organs will speak infallibly with respect the nature long lost structures species and groups species which are called aberrant and which may fancifully called living fossils will aid forming picture the ancient forms life embryology will reveal the structure some degree obscured the prototypes each great class when can feel assured that all the individuals the same species and all the closely allied species most genera have within not very remote period descended from one parent and have migrated from some one birthplace and when better know the many means migration then the light which geology now throws and will continue throw former changes climate and the level the land shall surely enabled trace admirable manner the former migrations the inhabitants the whole world even present comparing the differences the inhabitants the sea the opposite sides continent and the nature the various inhabitants that continent relation their apparent means immigration some light can thrown ancient geography the noble science geology loses glory from the extreme imperfection the record the crust the earth with its embedded remains must not looked well filled museum but poor collection made hazard and rare intervals the accumulation each great fossiliferous formation will recognised having depended unusual concurrence circumstances and the blank intervals between the successive stages having been vast duration but shall able gauge with some security the duration these intervals comparison the preceding and succeeding organic forms must cautious attempting correlate strictly contemporaneous two formations which include few identical species the general succession their forms life species are produced and exterminated slowly acting and still existing causes and not miraculous acts creation and catastrophes and the most important all causes organic change one which almost independent altered and perhaps suddenly altered physical conditions namely the mutual relation organism organism the improvement one being entailing the improvement the extermination others follows that the amount organic change the fossils consecutive formations probably serves fair measure the lapse actual time number species however keeping body might remain for long period unchanged whilst within this same period several these species migrating into new countries and coming into competition with foreign associates might become modified that must not overrate the accuracy organic change measure time during early periods the earth history when the forms life were probably fewer and simpler the rate change was probably slower and the first dawn life when very few forms the simplest structure existed the rate change may have been slow extreme degree the whole history the world present known although length quite incomprehensible will hereafter recognised mere fragment time compared with the ages which have elapsed since the first creature the progenitor innumerable extinct and living descendants was created the distant future see open fields for far more important researches psychology will based new foundation that the necessary acquirement each mental power and capacity gradation light will thrown the origin man and his history authors the highest eminence seem fully satisfied with the view that each species has been independently created mind accords better with what know the laws impressed matter the creator that the production and extinction the past and present inhabitants the world should have been due secondary causes like those determining the birth and death the individual when view all beings not special creations but the lineal descendants some few beings which lived long before the first bed the silurian system was deposited they seem become ennobled judging from the past may safely infer that not one living species will transmit its unaltered likeness distant futurity and the species now living very few will transmit progeny any kind far distant futurity for the manner which all organic beings are grouped shows that the greater number species each genus and all the species many genera have left descendants but have become utterly extinct can far take prophetic glance into futurity foretel that will the common and widely spread species belonging the larger and dominant groups which will ultimately prevail and procreate new and dominant species all the living forms life are the lineal descendants those which lived long before the silurian epoch may feel certain that the ordinary succession generation has never once been broken and that cataclysm has desolated the whole world hence may look with some confidence secure future equally inappreciable length and natural selection works solely and for the good each being all corporeal and mental endowments will tend progress towards perfection interesting contemplate entangled bank clothed with many plants many kinds with birds singing the bushes with various insects flitting about and with worms crawling through the damp earth and reflect that these elaborately constructed forms different from each other and dependent each other complex manner have all been produced laws acting around these laws taken the largest sense being growth with reproduction inheritance which almost implied reproduction variability from the indirect and direct action the external conditions life and from use and disuse ratio increase high lead struggle for life and consequence natural selection entailing divergence character and the extinction less improved forms thus from the war nature from famine and death the most exalted object which are capable conceiving namely the production the higher animals directly follows there grandeur this view life with its several powers having been originally breathed into few forms into one and that whilst this planet has gone cycling according the fixed law gravity from simple beginning endless forms most beautiful and most wonderful have been and are being evolved index aberrant groups 429 abyssinia plants 375 acclimatisation 139 affinities extinct species 329 organic beings 411 agassiz amblyopsis 139 groups species suddenly appearing 302 305 embryological succession 338 the glacial period 366 embryological characters 418 the embryos vertebrata 439 parallelism embryological development and geological succession 449 algæ new zealand 376 alligators males fighting amblyopsis blind fish 139 america north productions allied those europe 371 boulders and glaciers 373 south modern formations west coast 290 ammonites sudden extinction 321 anagallis sterility 247 analogy variations 159 ancylus 386 animals not domesticated from being variable domestic descended from several stocks acclimatisation 141 australia 116 with thicker fur cold climates 133 blind caves 137 extinct australia 339 anomma 240 antarctic islands ancient flora 399 antirrhinum 161 ants attending aphides 211 slave making instinct 219 ants neuter structure 236 aphides attended ants 211 aphis development 442 apteryx 182 arab horses aralo caspian sea 339 archiac the succession species 325 artichoke jerusalem 142 ascension plants 389 asclepias pollen 193 asparagus 359 aspicarpa 417 asses striped 163 ateuchus 135 audubon habits frigate bird 185 variation birds nests 212 heron eating seeds 387 australia animals 116 dogs 215 extinct animals 339 european plants 375 azara flies destroying cattle azores flora 363 babington british plants balancement growth 147 bamboo with hooks 197 barberry flowers barrande silurian colonies 313 the succession species 325 parallelism palæozoic formations 328 affinities ancient species 330 barriers importance 347 batrachians islands 393 bats how structure acquired 180 distribution 394 bear catching water insects 184 bee sting 202 queen killing rivals 202 bees fertilising flowers bees hive not sucking the red clover cell making instinct 224 humble cells 225 parasitic 218 beetles wingless madeira 135 with deficient tarsi 135 bentham british plants classification 419 berkeley seeds salt water 358 bermuda birds 391 birds acquiring fear 212 annually cross the atlantic 364 colour continents 132 fossil caves brazil 339 madeira bermuda and galapagos 390 song males transporting seeds 361 waders 386 wingless 134 182 with traces embryonic teeth 451 bizcacha 349 affinities 429 bladder for swimming fish 190 blindness cave animals 137 blyth distinctness indian cattle striped hemionus 163 crossed geese 253 boar shoulder pad borrow the spanish pointer bory vincent batrachians 393 bosquet fossil chthamalus 304 boulders erratic the azores 363 branchiæ 190 brent house tumblers 214 hawks killing pigeons 362 brewer american cuckoo 217 britain mammals 395 bronn duration specific forms 293 brown robert classification 414 buckman variation plants buzareingues sterility varieties 270 cabbage varieties crossed calceolaria 251 canary birds sterility hybrids 252 cape verde islands 398 cape good hope plants 110 375 carrier pigeons killed hawks 362 cassini flowers compositæ 145 catasetum 424 cats with blue eyes deaf variation habits curling tail when going spring 201 cattle destroying fir trees destroyed flies plata breeds locally extinct 111 fertility indian and european breeds 254 cave inhabitants blind 137 centres creation 352 cephalopodæ development 442 cervulus 253 cetacea teeth and hair 144 ceylon plants 375 chalk formation 322 characters divergence 111 sexual variable 156 adaptive analogical 427 charlock checks increase mutual chickens instinctive tameness 216 chthamalinæ 288 chthamalus cretacean species 304 circumstances favourable selection domestic products natural selection 101 cirripedes capable crossing 101 carapace aborted 148 their ovigerous frena 192 fossil 304 larvæ 440 classification 413 clift the succession types 339 climate effects checking increase beings adaptation organisms 139 cobites intestine 190 cockroach collections palæontological poor 287 colour influenced climate 132 relation attacks flies 198 columba livia parent domestic pigeons colymbetes 386 compensation growth 147 compositæ outer and inner florets 144 male flowers 451 conclusion general 480 conditions slight changes favourable fertility 267 coot 185 coral islands seeds drifted 360 reefs indicating movements earth 309 corn crake 185 correlation growth domestic productions growth 143 198 cowslip creation single centres 352 crinum 250 crosses reciprocal 258 crossing domestic animals importance altering breeds advantages unfavourable selection 102 crustacea new zealand 376 crustacean blind 137 cryptocerus 238 ctenomys blind 137 cuckoo instinct 216 currants grafts 262 currents sea rate 359 cuvier conditions existence 206 fossil monkeys 303 cuvier fred instinct 208 dana professor blind cave animals 139 relations crustaceans japan 372 crustaceans new zealand 376 candolle struggle for existence umbelliferæ 146 general affinities 430 candolle alph low plants widely dispersed 406 widely ranging plants being variable naturalisation 115 winged seeds 146 alpine species suddenly becoming rare 175 distribution plants with large seeds 360 vegetation australia 379 fresh water plants 386 insular plants 389 degradation coast rocks 282 denudation rate 285 oldest rocks 308 development ancient forms 336 devonian system 334 dianthus fertility crosses 256 dirt feet birds 362 dispersal means 356 during glacial period 365 distribution geographical 346 means 356 disuse effects under nature 134 divergence character 111 division physiological labour 115 dogs hairless with imperfect teeth descended from several wild stocks domestic instincts 213 inherited civilisation 215 fertility breeds together 254 crosses 268 proportions when young 444 domestication variation under downing fruit trees america downs north and south 285 dragon flies intestines 190 drift timber 360 driver ant 240 drones killed other bees 202 duck domestic wings reduced logger headed 182 duckweed 385 dugong affinities 414 dung beetles with deficient tarsi 135 dyticus 386 earl the malay archipelago 395 ears drooping domestic animals rudimentary 454 earth seeds roots trees 361 eciton 238 economy organisation 147 edentata teeth and hair 144 fossil species 339 edwards milne physiological divisions labour 115 gradations structure 194 embryological characters 418 eggs young birds escaping from electric organs 192 elephant rate increase glacial period 141 embryology 439 existence struggle for conditions 206 extinction bearing natural selection 109 domestic varieties 111 317 eye structure 187 correction for aberration 202 eyes reduced moles 137 fabre parasitic sphex 218 falconer naturalization plants india fossil crocodile 313 elephants and mastodons 334 and cautley mammals sub himalayan beds 340 falkland island wolf 393 faults 285 faunas marine 348 fear instinctive birds 212 feet birds young molluscs adhering 385 fertility hybrids 249 from slight changes conditions 267 crossed varieties 267 fir trees destroyed cattle pollen 203 fish flying 182 teleostean sudden appearance 305 eating seeds 362 387 fresh water distribution 384 fishes ganoid now confined fresh water 107 electric organs 192 ganoid living fresh water 321 southern hemisphere 376 flight powers how acquired 182 flowers structure relation crossing compositæ and umbelliferæ 144 forbes colours shells 132 abrupt range shells depth 175 poorness palæontological collections 287 continuous succession genera 316 continental extensions 357 distribution during glacial period 366 parallelism time and space 409 forests changes america formation devonian 334 formations thickness britain 284 intermittent 290 formica rufescens 219 formica sanguinea 219 formica flava neuter 239 frena ovigerous cirripedes 192 fresh water productions dispersal 383 fries species large genera being closely allied other species frigate bird 185 frogs islands 393 fruit trees gradual improvement united states varieties acclimatised united states 142 fuci crossed 258 fur thicker cold climates 133 furze 439 galapagos archipelago birds 390 productions 398 400 galeopithecus 181 game increase checked vermin gärtner sterility hybrids 247 255 reciprocal crosses 258 crossed maize and verbascum 270 comparison hybrids and mongrels 272 geese fertility when crossed 253 upland 185 genealogy important classification 425 geoffrey hilaire balancement 147 homologous organs 434 geoffrey hilaire isidore variability repeated parts 149 correlation monstrosities correlation 144 variable parts being often monstrous 155 geographical distribution 346 geography ancient 487 geology future progress 487 imperfection the record 279 giraffe tail 195 glacial period 365 gmelin distribution 365 gnathodon fossil 368 godwin austen the malay archipelago 299 goethe compensation growth 147 gooseberry grafts 262 gould land shells 397 gould colours birds 132 birds the galapagos 398 distribution genera birds 404 gourds crossed 270 grafts capacity 261 grasses varieties 113 gray asa trees united states 100 naturalised plants the united states 115 rarity intermediate varieties 176 alpine plants 365 gray striped mule 165 grebe 185 groups aberrant 429 grouse colours red doubtful species growth compensation 147 correlation domestic products correlation 143 habit effect under domestication effect under nature 134 diversified same species 183 hair and teeth correlated 144 harcourt the birds madeira 391 hartung boulders the azores 363 hazel nuts 359 hearne habits bears 184 heath changes vegetation heer plants madeira 107 helix pomatia 397 helosciadium 359 hemionus striped 163 herbert struggle for existence sterility hybrids 249 hermaphrodites crossing heron eating seed 387 heron sir peacocks heusinger white animals not poisoned certain plants hewitt sterility first crosses 264 himalaya glaciers 373 plants 375 hippeastrum 250 holly trees sexes hollyhock varieties crossed 271 hooker trees new zealand 100 hooker acclimatisation himalayan trees 140 flowers umbelliferæ 145 glaciers himalaya 373 algæ new zealand 376 vegetation the base the himalaya 378 plants tierra del fuego 374 378 australian plants 375 399 relations flora south america 379 flora the antarctic lands 381 399 the plants the galapagos 391 398 hooks bamboos 197 seeds islands 392 horner the antiquity egyptians horns rudimentary 454 horse fossil plata 318 horses destroyed flies plata striped 163 proportions when young 445 horticulturists selection applied huber cells bees 230 huber reason blended with instinct 208 habitual nature instincts 208 slave making ants 219 melipona domestica 225 humble bees cells 225 hunter secondary sexual characters 150 hutton captain crossed geese 253 huxley professor structure hermaphrodites 101 embryological succession 338 homologous organs 438 the development aphis 442 hybrids and mongrels compared 272 hybridism 245 hydra structure 190 ibla 148 icebergs transporting seeds 363 increase rate individuals numbers favourable selection 102 many whether simultaneously created 356 inheritance laws corresponding ages insects colour fitted for habitations sea side colours 132 blind caves 138 luminous 193 neuter 236 instinct 207 instincts domestic 213 intercrossing advantages islands oceanic 388 isolation favourable selection 104 japan productions 372 java plants 375 jones the birds bermuda 391 jussieu classification 417 kentucky caves 137 kerguelen land flora 381 399 kidney bean acclimatisation 142 kidneys birds 144 kirby tarsi deficient beetles 135 knight andrew cause variation kölreuter the barberry sterility hybrids 247 reciprocal crosses 258 crossed varieties nicotiana 271 crossing male and hermaphrodite flowers 451 lamarck adaptive characters 427 land shells distribution 397 madeira naturalised 402 languages classification 422 lapse great time 282 larvæ 440 laurel nectar secreted the leaves laws variation 131 leech varieties leguminosæ nectar secreted glands lepidosiren 107 330 life struggle for lingula silurian 306 linnæus aphorism 413 lion mane young striped 439 lobelia fulgens lobelia sterility crosses 250 loess the rhine 384 lowness structure connected with variability 149 lowness related wide distribution 406 lubbock the nerves coccus lucas inheritance resemblance child parent 275 lund and clausen fossils brazil 339 lyell sir the struggle for existence modern changes the earth measure denudation 283 carboniferous land shell 289 fossil whales 303 strata beneath silurian system 307 the imperfection the geological record 310 the appearance species 312 barrande colonies 313 tertiary formations europe and north america 323 parallelism tertiary formations 328 transport seeds icebergs 363 great alternations climate 382 the distribution fresh water shells 385 land shells madeira 402 lyell and dawson fossilized trees nova scotia 296 macleay analogical characters 427 madeira plants 107 beetles wingless 135 fossil land shells 339 birds 390 magpie tame norway 212 maize crossed 270 malay archipelago compared with europe 299 mammals 395 malpighiaceæ 417 mammæ rudimentary 451 mammals fossil secondary formation 303 insular 393 man origin races 199 manatee rudimentary nails 454 marsupials australia 116 fossil species 339 martens experiment seeds 360 martin striped mules 165 matteuchi the electric organs rays 193 matthiola reciprocal crosses 258 means dispersal 356 melipona domestica 225 metamorphism oldest rocks 308 mice destroying bees acclimatisation 141 migration bears first appearance fossils 296 miller professor the cells bees 226 mirabilis crosses 258 missel thrush misseltoe complex relations mississippi rate deposition mouth 284 mocking thrush the galapagos 402 modification species how far applicable 483 moles blind 137 mongrels fertility and sterility 267 and hybrids compared 272 monkeys fossil 303 monocanthus 424 mons van the origin fruit trees moquin tandon sea side plants 132 morphology 434 mozart musical powers 209 mud seeds 386 mules striped 165 müller alpine australian plants 375 murchison sir the formations russia 289 azoic formations 307 extinction 317 mustela vison 179 myanthus 424 myrmecocystus 238 myrmica eyes 240 nails rudimentary 453 natural history future progress 484 selection system 413 naturalisation forms distinct from the indigenous species 115 new zealand 201 nautilus silurian 306 nectar plants nectaries how formed nelumbium luteum 387 nests variation 212 neuter insects 236 newman humble bees new zealand productions not perfect 201 naturalised products 337 fossil birds 339 glacial action 373 crustaceans 376 algæ 376 number plants 389 flora 399 nicotiana crossed varieties 271 certain species very sterile 257 noble fertility rhododendron 251 nodules phosphatic azoic rocks 307 oak varieties onites apelles 135 orchis pollen 193 organs extreme perfection 186 electric fishes 192 little importance 194 homologous 434 rudiments 450 ornithorhynchus 107 416 ostrich not capable flight 134 habit laying eggs together 218 american two species 349 otter habits how acquired 179 ouzel water 185 owen professor birds not flying 134 vegetative repetition 149 variable length arms ourang outang 150 the swim bladder fishes 191 electric organs 192 fossil horse plata 319 relations ruminants and pachyderms 329 fossil birds new zealand 339 succession types 339 affinities the dugong 414 homologous organs 435 the metamorphosis cephalopods and spiders 442 pacific ocean faunas 348 paley organ formed give pain 201 pallas the fertility the wild stocks domestic animals 253 paraguay cattle destroyed flies parasites 217 partridge dirt feet 362 parts greatly developed variable 150 degrees utility 201 parus major 183 passiflora 251 peaches united states pear grafts 261 pelargonium flowers 145 sterility 251 pelvis women 144 peloria 145 period glacial 365 petrels habits 184 phasianus fertility hybrids 253 pheasant young wild 216 philippi tertiary species sicily 312 pictet professor groups species suddenly appearing 302 305 rate organic change 313 continuous succession genera 316 close alliance fossils consecutive formations 335 embryological succession 338 pierce varieties wolves pigeons with feathered feet and skin between toes breeds described and origin breeds how produced tumbler not being able get out egg reverting blue colour 160 instinct tumbling 214 carriers killed hawks 362 young 445 pistil rudimentary 451 plants poisonous not affecting certain coloured animals selection applied gradual improvement not improved barbarous countries destroyed insects midst range have struggle with other plants nectar fleshy sea shores 132 fresh water distribution 386 low scale widely distributed 406 plumage laws change sexes birds plums the united states pointer dog origin habits 213 poison not affecting certain coloured animals poison similar effect animals and plants 484 pollen fir trees 203 poole col striped hemionus 163 potamogeton 387 prestwich english and french eocene formations 328 primrose sterility 247 primula varieties proteolepas 148 proteus 139 psychology future progress 488 quagga striped 165 quince grafts 261 rabbit disposition young 215 races domestic characters race horses arab english 356 ramond plants pyrenees 368 ramsay professor thickness the british formations 284 faults 285 ratio increase rats supplanting each other acclimatisation 141 blind cave 137 rattle snake 201 reason and instinct 208 recapitulation general 459 reciprocity crosses 258 record geological imperfect 279 rengger flies destroying cattle reproduction rate resemblance parents mongrels and hybrids 273 reversion law inheritance pigeons blue colour 160 rhododendron sterility 251 richard professor aspicarpa 417 richardson sir structure squirrels 180 fishes the southern hemisphere 376 robinia grafts 262 rodents blind 137 rudimentary organs 450 rudiments important for classification 416 sageret grafts 262 salmons males fighting and hooked jaws salt water how far injurious seeds 358 saurophagus sulphuratus 183 schiödte blind insects 138 schlegel snakes 144 sea water how far injurious seeds 358 sebright sir crossed animals selection pigeons sedgwick professor groups species suddenly appearing 302 seedlings destroyed insects seeds nutriment winged 146 power resisting salt water 358 crops and intestines birds 361 eaten fish 362 387 mud 386 hooked islands 392 selection domestic products principle not recent origin unconscious natural sexual natural circumstances favourable 101 sexes relations sexual characters variable 156 selection sheep merino their selection two sub breeds unintentionally produced mountain varieties shells colours 132 littoral seldom embedded 288 fresh water dispersal 385 madeira 391 land distribution 397 silene fertility crosses 257 silliman professor blind rat 137 skulls young mammals 197 437 slave making instinct 219 smith col hamilton striped horses 164 smith fred slave making ants 219 neuter ants 239 smith jordan hill the degradation coast rocks 283 snap dragon 161 somerville lord selection sheep sorbus grafts 262 spaniel king charles breed species polymorphic common variable large genera variable groups suddenly appearing 302 306 beneath silurian formations 306 successively appearing 312 changing simultaneously throughout the world 322 spencer lord increase size cattle sphex parasitic 218 spiders development 442 spitz dog crossed with fox 268 sports plants sprengel crossing ray florets 145 squirrels gradations structure 180 staffordshire heath changes stag beetles fighting sterility from changed conditions life hybrids 246 laws 254 causes 263 from unfavourable conditions 265 certain varieties 269 helena productions 389 hilaire aug classification 418 john habits cats sting bee 202 stocks aboriginal domestic animals strata thickness britain 284 stripes horses 163 structure degrees utility 201 struggle for existence succession geological 312 succession types same areas 338 swallow one species supplanting another swim bladder 190 system natural 413 tail giraffe 195 aquatic animals 196 rudimentary 454 tarsi deficient 135 tausch umbelliferous flowers 146 teeth and hair correlated 144 embryonic traces birds 451 rudimentary embryonic calf 450 480 tegetmeier cells bees 228 233 temminck distribution aiding classification 419 thouin grafts 262 thrush aquatic species 185 mocking the galapagos 402 young spotted 439 nest 243 thuret crossed fuci 258 thwaites acclimatisation 140 tierra del fuego dogs 215 plants 374 378 timber drift 360 time lapse 282 titmouse 183 toads islands 393 tobacco crossed varieties 271 tomes the distribution bats 394 transitions varieties rare 172 trees islands belong peculiar orders 392 with separated sexes trifolium pratense trifolium incarnatum trigonia 321 trilobites 306 sudden extinction 321 troglodytes 243 tucutucu blind 137 tumbler pigeons habits hereditary 214 young 446 turkey cock brush hair breast turkey naked skin head 197 young wild 216 turnip and cabbage analogous variations 159 type unity 206 types succession same areas 338 udders enlarged use rudimentary 451 ulex young leaves 439 umbelliferæ outer and inner florets 144 unity type 206 use effects under domestication effects state nature 134 utility how far important the construction each part 199 valenciennes fresh water fish 384 variability mongrels and hybrids 274 variation under domestication caused reproductive system being affected conditions life under nature laws 131 variations appear corresponding ages analogous distinct species 159 varieties natural struggle between domestic extinction 111 transitional rarity 172 when crossed fertile 267 when crossed sterile 269 classification 423 verbascum sterility 251 varieties crossed 270 verneuil the succession species 325 viola tricolor volcanic islands denudation 284 vulture naked skin head 197 wading birds 386 wallace origin species law geographical distribution 355 the malay archipelago 395 wasp sting 202 water fresh productions 383 water hen 185 waterhouse australian marsupials 116 greatly developed parts being variable 150 the cells bees 225 general affinities 429 water ouzel 185 watson range varieties british plants acclimatisation 140 flora azores 363 alpine plants 367 376 rarity intermediate varieties 176 weald denudation 285 web feet water birds 185 west indian islands mammals 395 westwood species large genera being closely allied others the tarsi engidæ 157 the antennæ hymenopterous insects 416 whales fossil 303 wheat varieties 113 white mountains flora 365 wings reduction size 134 wings insects homologous with branchiæ 191 rudimentary insects 451 wolf crossed with dog 214 falkland isles 393 wollaston varieties insects fossil varieties land shells madeira colours insects sea shore 132 wingless beetles 135 rarity intermediate varieties 176 insular insects 389 land shells madeira naturalised 402 wolves varieties woodpecker habits 184 green colour 197 woodward the duration specific forms 293 the continuous succession genera 316 the succession types 339 world species changing simultaneously throughout 322 wrens nest 243 youatt selection sub breeds sheep rudimentary horns young cattle 454 zebra stripes 163 end the project gutenberg ebook the origin species means natural selection updated editions will replace the previous one the old editions will renamed creating the works from print editions not protected copyright law means that one owns united states copyright these works the foundation and you can copy and distribute the united states without permission and without paying copyright royalties special rules 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